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CYPERACEAE

SEDGE FAMILY

Raymond Cranfill, except as specified

Annual or perennial herb, often rhizomed, often of wet open places, generally monoecious; roots fibrous, hairy
Stem generally 3-sided
Leaves often 3-ranked; sheath generally closed; ligule generally 0; blade (0) various, parallel-veined
Inflorescence: spikelets variously clustered; flowers generally sessile in axil of flower bract
Flower small, generally wind-pollinated; perianth 0 or bristle-like; stamens generally 3, anthers attached at base, 4-chambered; ovary superior, 1-chambered, 1-ovuled, style 2–3-branched
Fruit: achene, generally 3-sided
Genera in family: ± 110 genera, 3600 species: worldwide, especially temp
Reference: [Tucker 1987 J Arnold Arbor 68:361–445]
Difficult: taxa differ in technical characters of inflorescence and fruit.

ELEOCHARIS

SPIKERUSH

Annual or perennial herb
Stem generally round, ridged and grooved, generally solid
Leaves basal, 1(–4); base sheathing; blade generally 0
Inflorescence: spikelet solitary, terminal, erect; spikelet bract 0; flower bracts generally spiraled
Flowers bisexual; perianth bristles 0–6, persistent, barbs generally recurved; stamens (1–)3; style 2–3-branched, base bulb-like, persistent
Fruit 2–3-sided or round; top tubercled
Species in genus: ± 250 species: worldwide
Etymology: (Greek: marsh grace)
St shape best seen in fresh material (or just below spikelet); drying exaggerates ridges and grooves.

Native

E. atropurpurea (Retz.) Kunth

Annual < 1.2 dm
Leaf: base deep brown; tip oblique, 1-toothed, green to reddish
Inflorescence: spikelet 3–8 mm, much wider than stem, oblong-ovate, generally 10–many-flowered; flower bracts ovate-obtuse, greenish with dark brown sides, lower deciduous
Flower: style 2-branched
Fruit: perianth bristles generally < fruit; body ± 0.5 mm, obovate, sharply 2-sided, black, shiny; tubercle < 1/5 fruit body, hat-like, base strongly narrowed
Ecology: Rice fields, wet fallow ground
Elevation: < 500 m.
Bioregional distribution: South Coast, Great Central Valley
Distribution outside California: to Washington, c&se US; tropical

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