Jepson Flora Project: Jepson Interchange    

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Jepson Interchange (more information)
©Copyright 1993 by the Regents of the University of California

  • Up-to-date information about California vascular plants is available from the Jepson eFlora.



Patrick E. Elvander

Perennial or subshrub from caudex or rhizome, generally ± hairy
Stem often ± leafy on lower half, rarely trailing and leafy throughout
Leaves generally simple, basal or sometimes cauline, generally alternate, generally petioled; veins ± palmate
Inflorescence: panicle, generally ± scapose
Flower generally bisexual, generally radial; hypanthium free to ± fused to ovary; calyx lobes generally 5; petals generally 5, free, generally clawed, generally white; stamens generally 5 or 10; pistils 2 and simple or 1 and compound (chambers 1–2, placentas 2–4, axile or parietal), ovary superior to inferior, sometimes more superior in fruit, styles generally 2
Fruit: 2 follicles or 2–4-valved capsule
Seeds generally many, small
Genera in family: 40 genera, 600 species: especially n temp, arctic, alpine; some cultivated (Bergenia , Darmera , Heuchera , Saxifraga , Tellima , Tolmiea )
Reference: [Soltis 1988 Syst Bot 13:64–72]



Plant generally ± hairy, often glandular; caudex or rhizome generally not woody, generally scaly
Leaf: blade oblanceolate to round, base tapered to reniform, margin entire or toothed
Inflorescence: flowers 1–many; bracts scale-like
Flower: hypanthium free or ± fused to ovary; petals sometimes spotted; stamens 10, filaments generally flat; pistil 1 (chambers 2, placentas 2, axile) or 2 (each with 1 chamber, 1 marginal placenta), ovary superior to ± inferior (sometimes more superior in fruit), styles 2, free throughout
Fruit: capsule or 2 follicles
Species in genus: ± 400 species: especially cool n temp
Etymology: (Latin: rock-breaking)
Reference: [Elvander 1984 Syst Bot Monogr 3:1–44]


S. marshallii Greene

Caudex generally producing rhizomes; bulblets 0
Leaf 5–15 cm; petiole 3–10 cm; blade ovate or elliptic, base truncate to tapered, teeth coarse, sharp to round
Inflorescence 10–40 cm, open
Flower: sepals reflexed, generally < petals, ovate to elliptic; petals 2.5–4.5 mm, elliptic or obovate, 2-spotted, ephemeral; filaments generally much-widened above middle; nectaries band-like; ovary superior
Fruit: ± follicles
Ecology: Mossy rocks, cliffs
Elevation: < 1500 m.
Bioregional distribution: North Coast, Klamath Ranges, Outer North Coast Ranges
Distribution outside California: sw Oregon

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