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SALICACEAE

WILLOW FAMILY

Shrub, tree, generally dioecious (rarely monoecious)
Stem: trunk < 40 m; wood soft; bark smooth, bitter; buds scaly
Leaves simple, alternate, deciduous; stipules generally deciduous, often large
Inflorescence: catkin, generally appearing before leaves; each flower subtended by disk or 1–2 nectary glands and 1 bract
Flower: perianth 0
Staminate flower: stamens 1–many
Pistillate flower: pistil 1, ovary superior, chamber 1, stigma lobes 2–4
Seeds many; hairs fine, white, cottony
Fruit: capsule; valves 2–4
Genera in family: 2 genera, 340 species: generally temp (except Australia, Malay Archipelago) moist places; many cultivated. Hybridization common; identification often difficult. Family description, key to genera by John O. Sawyer, Jr.

SALIX

WILLOW

George W. Argus

Shrub, tree, dioecious; bud scale 1, not sticky, margins generally fused (or free, overlapping)
Stem: twigs generally flexible and not glaucous
Leaf: blade linear to widely obovate, entire to toothed, generally ± hairy
Inflorescence: dense catkin emerging before, with, or after leaves, sessile or on a short leafy shoot; bract subtending each flower
Flower: perianth 0
Staminate flower: stamens 1–8
Pistillate flower: ovary stalked or sessile, style 1 or 0, stigmas 2, each sometimes 2-lobed; nectaries 1–several, generally rod-like, generally between inflorescence axis and flower
Fruit: valves 2
Species in genus: ± 400 species: ± worldwide, especially n temp, arctic
Etymology: (Latin: ancient name)
Reference: [Argus 1986 Syst Bot Monog 9:1–170; Dorn 1976 Canad J Bot 54:2769–2789]
Difficult, highly variable. Not all specimens will key easily; sprout shoots and other extreme forms are not included in keys, may require field comparison for identification. Studies of variation, hybridization needed.

Native

S. sessilifolia Nutt.

SANDBAR WILLOW

Shrub < 5 m, clonal by root-sprouting
Stem: twigs brownish, persistently silky, long-soft-wavy-hairy, or becoming glabrous
Leaves: petiole very short; young leaves hairy; mature blade 40–120 mm, ± elliptic to narrowly obovate, acuminate (base wedge-shaped), sparsely spiny-serrate, silky, lower surface not glaucous
Inflorescence appearing with or after leaves, branched, 20–70 mm, on leafy shoots 20–125 mm; flower bract tawny or brown, tip rounded or acute; pistillate bract deciduous after flower
Staminate flower: stamens 2
Pistillate flower: ovary long-soft-wavy-hairy, stigmas deciduous after flower, style 0.2–0.4 mm, stalk 0.2–0.7 mm
Ecology: Streambanks, sandbars
Elevation: generally < 200 m.
Bioregional distribution: North Coast, w Klamath Ranges
Distribution outside California: to British Columbia
Synonyms: S. parksiana C.R. Ball, Del Norte coast willow
Horticultural information: WET: 4, 5, 6, 7, 14, 15, 16, 17, 19, 20, 21, 22, 23, 24; STBL.

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