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©Copyright 1993 by the Regents of the University of California
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ROSACEAE

ROSE FAMILY

Annual to tree
Leaves simple to pinnately to palmately compound, generally alternate; stipules free to fused, persistent to deciduous
Inflorescence: cyme, raceme, panicle, or flowers solitary
Flower generally bisexual, radial; hypanthium free or fused to ovary, saucer- to funnel-shaped, often with bractlets alternate with sepals; sepals generally 5; petals generally 5, free; stamens (0)5–many, pistils (0)1–many, simple or compound; ovary superior to inferior, styles 1–5
Fruit: achene, follicle, drupe, pome, or blackberry- to raspberry-like
Seeds generally 1–5
Genera in family: 110 genera, ± 3000 species: worldwide, especially temp. Many cultivated for ornamental and fruit, especially Cotoneaster , Fragaria , Malus , Prunus , Pyracantha, Rosa , and Rubus
Reference: [Robertson 1974 J Arnold Arbor 55:303–332,344–401,611–662]
Family description, key to genera by Barbara Ertter and Dieter H. Wilken.

CRATAEGUS

HAWTHORN

Thomas J. Rosatti

Shrub or tree, generally thorny
Leaves stipuled, petioled, deciduous, toothed to lobed
Inflorescence: racemes at branch tips
Flower: hypanthium cup- to urn-shaped, bractlets 0; sepals becoming reflexed; petals generally white; stamens 5–25; ovary inferior, styles 1–5, free
Fruit: pome, drupe-like, purple-black and glabrous in CA; stones 1–5, generally 1-seeded
Species in genus: ± 300 species: n temp
Etymology: (Greek: flower thorn, used by Theophrastus?; or strength, from strong wood or thorns)
Reference: [Brunsfeld & Johnson 1990 Madroño 37:274–282]
C. erythropoda may occur in e D; C. monogyna Jacq. escaped from or persistent beyond cultivated, perhaps naturalized. Many hybrids and morphologically identifiable clones.

Native

C. suksdorfii (Sarg.) Kruschke


Stem: older thorns 8–12 mm
Leaf 2–9 cm; blade base wedge-shaped, entire, margin above base toothed, generally unlobed, tips rounded
Flower 12–13 mm wide when dry; sepals entire to ± gland-toothed; stamens (12–)20; ovary generally glabrous, styles generally 5
Fruit generally few per inflorescence, 7–8 mm wide when fresh; stone ± 4.5 mm, outer face keeled, narrow, sides deeply pitted
Chromosomes: 2n=34
Ecology: Moist streamsides, lakesides, forest
Elevation: generally < 1500 m.
Bioregional distribution: Klamath Ranges, North Coast Ranges, Cascade Range, n San Francisco Bay Area
Distribution outside California: to British Columbia, Montana
Horticultural information: 4, 5, 6 &IRR: 1, 2, 3, 7, 8, 9, 14, 15, 16, 17, 18, 19, 20, 21, 22, 23, 24; may be INV.

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