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RANUNCULACEAE

BUTTERCUP FAMILY

Dieter H. Wilken, except as specified

Annual, perennial herb, sometimes aquatic
Leaves generally basal and cauline, generally alternate, simple or compound; petioles at base generally flat, sometimes sheathing or stipule-like
Inflorescence: cyme, raceme, panicle, or flowers solitary
Flower generally bisexual, radial; sepals generally 5, free, early deciduous or withering in fruit, generally green; petals 0–many, free; stamens generally 10–many; pistils 1–many, ovary superior, chamber 1, style 1, generally ± persistent in fruit as beak, ovules 1–many
Fruit: achene, follicle, berry, or utricle-like, 1–many-seeded
Genera in family: ± 60 genera, 1700 species: worldwide, especially n temp, tropical mtns; many ornamental (Adonis, Aquilegia, Clematis, Consolida, Delphinium, Erianthis, Helleborus ), some highly TOXIC (Aconitum, Actaea, Delphinium, Ranunculus )
Reference: [Duncan & Keener 1991 Phytologia 70:24–27]

KUMLIENIA

Perennial from short, stout caudex, generally ± glabrous; roots clustered, fleshy
Leaves simple, basal, rosetted
Inflorescence scapose, 1(–3)-flowered; peduncle long
Flower bisexual, radial; sepals 5–6, petal-like; petals 5–12, < sepals, gland-like; pistils many, style persistent in fruit, beak-like
Fruit utricle-like, splitting tardily from base along 2 lines; wall thin, slightly transparent between veins; beak tapered, straight, tip hooked
Seed 1, cylindric
Species in genus: 2 species: w North America
Etymology: (T.L. Kumlien, Swedish naturalist, mentor of E.L. Greene, 19th century)
Reference: [Greene 1886 Bull Calif Acad Sci 1:337–338]

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