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  • Up-to-date information about California vascular plants is available from the Jepson eFlora.

ASTERACEAE

SUNFLOWER FAMILY

David J. Keil, Family Editor and author, except as specified

Annual to tree
Leaves basal or cauline, alternate to whorled, simple to compound
Inflorescence: 1° inflorescence a head, each resembling a flower, 1–many, generally arrayed in cymes, generally subtended by ± calyx-like involucre; flowers 1–many per head
Flowers bisexual, unisexual, or sterile, ± small, of several types; calyx 0 or modified into pappus of bristles, scales, or awns, which is generally persistent in fruit; corolla radial or bilateral (rarely 0), lobes generally (0)4–5; stamens 4–5, anthers generally fused into cylinder around style, often appendaged at tips, bases, or both, filaments generally free, generally attached to corolla near throat; pistil 1, ovary inferior, 1-chambered, 1-seeded, style 1, branches 2, generally hair-tufted at tip, stigmas 2, generally on inside of style branches
Fruit: achene, cylindric to ovoid, generally deciduous with pappus attached
Genera in family: ± 1300 genera, 21,000 species (largest family of dicots): worldwide. Largest family in CA. Also see tribal key to CA genera: Strother 1997 Madroño 44(1):1–28. See glossary p. 25 for illustrations of general family characteristics.

CENTAUREA

KNAPWEED, STAR-THISTLE

David J. Keil and Charles E. Turner

Annual to perennial herb, ± branched
Leaves alternate; lower generally deeply 1–2-lobed, segments generally narrow; upper reduced
Inflorescence: heads discoid (sterile outer flowers sometimes ± ray-like); involucre cylindric to hemispheric; phyllaries many, graded, generally ± ovate, scarious-margined, tip appendages fringed to spiny; receptacle flat, long-bristly
Flowers: inner fruiting; anther bases tailed, tips oblong; style top minutely hairy, tips minutely branched
Fruit ± barrel-shaped, ± compressed, attached ± at side; pappus generally of stiff, unequal bristles or narrow scales
Species in genus: ± 500 species: especially Eurasia, n Africa (± 2 North America); some cultivated (waifs may including C. cineraria L., C. eriophora L., C. jacea L., C. moschata L., C. muricata L., C. salmantica L.)
Etymology: (Greek: ancient name)
Weedy.

Introduced

C. maculosa Lam.

SPOTTED KNAPWEED

Biennial 3–10 dm, ± gray-tomentose
Leaves resin-dotted; lower 10–15 cm, ± deeply 1–2-lobed
Inflorescence open; heads generally many; involucre 10–13 mm, ovoid; phyllaries pale green or pink-tinged, prominently parallel-veined, appendages fringed with slender, dark teeth
Flowers 30–40; corollas 12–25 mm, white to pink or purple, sterile corollas ± slender
Fruit 3–3.5 mm, ± pale brown, finely hairy; pappus bristles 1–2 mm, white
Chromosomes: 2n=18,36
Ecology: Disturbed areas
Elevation: < 2000 m.
Bioregional distribution: Northwestern California, Cascade Range, Sierra Nevada, n Sacramento Valley, n Central Western California, s Peninsular Ranges, Modoc Plateau, n East of Sierra Nevada
Distribution outside California: native to Europe
Flowering time: Summer
Weedy.

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bioregional map for CENTAUREA%20maculosa being generated
 
N.B. The distribution depicted here differs from that given in The Jepson Manual (1993)

Retrieve Jepson Interchange Index to Plant Names entry for Centaurea maculosa
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