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Jepson Interchange (more information)
©Copyright 1993 by the Regents of the University of California

  • Up-to-date information about California vascular plants is available from the Jepson eFlora.



David J. Keil, Family Editor and author, except as specified

Annual to tree
Leaves basal or cauline, alternate to whorled, simple to compound
Inflorescence: 1° inflorescence a head, each resembling a flower, 1–many, generally arrayed in cymes, generally subtended by ± calyx-like involucre; flowers 1–many per head
Flowers bisexual, unisexual, or sterile, ± small, of several types; calyx 0 or modified into pappus of bristles, scales, or awns, which is generally persistent in fruit; corolla radial or bilateral (rarely 0), lobes generally (0)4–5; stamens 4–5, anthers generally fused into cylinder around style, often appendaged at tips, bases, or both, filaments generally free, generally attached to corolla near throat; pistil 1, ovary inferior, 1-chambered, 1-seeded, style 1, branches 2, generally hair-tufted at tip, stigmas 2, generally on inside of style branches
Fruit: achene, cylindric to ovoid, generally deciduous with pappus attached
Genera in family: ± 1300 genera, 21,000 species (largest family of dicots): worldwide. Largest family in CA. Also see tribal key to CA genera: Strother 1997 Madroño 44(1):1–28. See glossary p. 25 for illustrations of general family characteristics.


Annual, perennial herb
Stems slender; base ± leafy
Leaves alternate, reduced upward; lower leaves long-petioled; blades wide, thin
Inflorescence: heads small, disciform, in open panicle-like cymes; involucre inconspicuous; phyllaries in 1–2 series, ± equal, reflexed in fruit, deciduous; receptacle convex, naked
Pistillate flowers few in 1 series; corolla deeply 4–5-lobed, white, soon deciduous
Disk flowers few, staminate, ± persistent; corolla cylindric, white; anther bases sagittate, tips narrowly triangular; style undivided
Fruit club-shaped to obovoid, weakly veined, prominently stalked-glandular; pappus 0
Species in genus: 6 species: w North America, C.Am, s South America, e Asia
Etymology: (Greek: gland stem)


A. bicolor Hook.


Perennial 3–10 dm, generally ± erect, openly branched, ± tomentose below, stalked-glandular above
Leaves at base; petiole ± winged; blade 3–25 cm, ± triangular to ovate, base cordate or hastate, subentire to shallowly lobed, white-tomentose below, glabrous above
Inflorescence: phyllaries 1–2.5 mm, ovate
Pistillate flowers 3–7; corollas 0.5–1 mm
Disk flowers 3–10; corollas 1.5–2 mm
Fruit 5–8 mm, club-shaped
Chromosomes: 2n=46
Ecology: Generally in shade, woods, forests
Elevation: < 2000 m.
Bioregional distribution: Northwestern California, Cascade Range, Sierra Nevada, nw Central Western California
Distribution outside California: to British Columbia, north-central US

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