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Warren L. Wagner, except as specified Peter H. Raven, Family Coordinator

Annual to tree
Leaves basal or cauline, alternate, opposite, or whorled, generally simple and toothed (to pinnately compound); stipules 0 or generally deciduous
Inflorescence: spike, raceme, panicle, or flowers solitary in axils; bracted
Flower generally bisexual, generally radial, opening at dawn or dusk; hypanthium sometimes prolonged beyond ovary (measured from ovary tip to sepal base); sepals generally 4(2–7); petals generally 4 (or as many as sepals, rarely 0), often "fading" darker; stamens generally 4 or 8(2), anthers 2-chambered, opening lengthwise, pollen generally interconnected by threads; ovary inferior, chambers generally 4 (sometimes becoming 1), placentas axile or parietal, ovules 1–many per chamber, style 1, stigma 4-lobed (or lobes as many as sepals), club-shaped, or hemispheric
Fruit: capsule, loculicidal (sometimes berry or indehiscent and nut-like)
Seeds sometimes winged or hair-tufted
Genera in family: 15 genera, ± 650 species: worldwide, especially w North America; many cultivated (Clarkia, Epilobium, Fuchsia, Gaura, Oenothera )
Reference: [Munz 1965 North America Fl II 5:1–278]


Harlan Lewis

Stem prostrate to erect, < 1.5 m, glabrous, often glaucous, or puberulent; hairs rarely long and spreading
Leaf simple, pinnately veined; petiole < 4 cm or 0; blade 1–10 cm, linear to elliptic or ovate, entire or shallow-toothed, glabrous or sparsely puberulent
Inflorescence: spike, raceme; bracts leaf-like; axis in bud straight or recurved at tip, in flower ± straight; buds erect or not
Flower: hypanthium obconic to cup-shaped or long, slender, generally with ring of hairs within; sepals 4, generally fused to tip in bud, reflexed at least at base, staying fused at least at tip, in 4's or 2's, or all coming free; corolla bowl-shaped to rotate, petals 5–60 mm, often lobed or clawed, lavender or pink to dark red, pale yellow, or white, often spotted, flecked, or streaked with red, purple, or white; stamens 8, in 2 like or unlike series, or 4, filaments cylindric to wider above, subtended by ciliate scales or generally not, anthers attached at base, pollen white or yellow to blue-gray, lavender, or reddish; ovary 4-chambered, glabrous or not, cylindric, fusiform, or wider above, generally shallowly to deeply 4- or 8-grooved, stigma lobes 4, generally prominent
Fruit: generally capsule, elongate, rarely short, indehiscent, nut-like
Seeds generally many, rarely 1–2, 0.5–2 mm, angled, crested or not, brown, gray or mottled
Species in genus: ± 41 species: w North America, 1 South America
Etymology: (Captain William Clark, 1770–1838, of Lewis & Clark Expedition)
Reference: [Lewis & Lewis 1955 UC Publ Bot 20:241–392]
Self-compatible or -pollinated or outcrossed; on herbarium specimens, curvature of inflorescence axis in bud generally reliable, pollen color generally not.

Information on Stem description was contributed by Bob Allen (15 Aug 2004):
I think that should read prostRate, not prostate


C. tembloriensis Vasek

Stem erect, < 8 dm
Leaf: petiole 0–5 mm; blade 2–7 cm, lanceolate, glaucous, gray-green
Inflorescence: axis in bud straight; buds reflexed
Flower: hypanthium 2–3 mm; sepals staying fused in 4's, green, red-tinged or not, sparsely to densely puberulent; corolla rotate, petals 1–2.5 cm, lavender-pink, claw ± = or > blade, slender, blade ± diamond-shaped, spot purplish or 0; stamens 8, outer anthers lavender to red, inner smaller, paler; ovary 8- grooved, hairs as on sepals, stigma beyond anthers or not
Fruit 1.5–3 cm, ± 3 mm wide
Ecology: Dry grassland, scrub
Elevation: 100–500 m.
Bioregional distribution: s Sierra Nevada Foothills, San Joaquin Valley, Inner South Coast Ranges. Hybrids between subspp. have low fertility.


subsp. tembloriensis

Flower: petals generally < 10 mm wide; stigma beyond anthers or not
Chromosomes: n=9
Ecology: Habitats, elevations, range of sp
Bioregional distribution: s Sierra Nevada Foothills, San Joaquin Valley, Inner South Coast Ranges
Horticultural information: TRY.

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bioregional map for CLARKIA%20tembloriensis%20subsp.%20tembloriensis being generated

Retrieve Jepson Interchange Index to Plant Names entry for Clarkia tembloriensis subsp. tembloriensis
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