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©Copyright 1993 by the Regents of the University of California

  • Up-to-date information about California vascular plants is available from the Jepson eFlora.



Dieter H. Wilken, except as specifed

Annual, perennial herb, shrub, glabrous to hairy, generally aromatic
Stems generally erect, generally 4-angled
Leaves generally simple to deeply lobed, opposite, generally gland-dotted
Inflorescence: cyme, generally clustered around stem, head-like, separated by evident internodes (terminal in Monardella ) or collectively crowded, spike-like to panicle-like (sometimes raceme or flowers 2–12); subtended by leaves or bracts; flowers sessile or pedicelled
Flower generally bisexual; calyx generally 5-lobed, radial to bilateral; corolla generally bilateral, 1–2-lipped, upper lip entire or 2-lobed, ± flat to hood-like, sometimes 0, lower lip generally 3-lobed; stamens generally 4, generally exserted, paired, pairs unequal, sometimes 2, staminodes 2 or 0; ovary superior, generally 4-lobed to base, chambers 2, ovules 2 per chamber, style 1, arising from center at junction of lobes, stigmas generally 2
Fruit: nutlets 4, generally ovoid to oblong, smooth
Genera in family: ± 200 genera, 5500 species: worldwide. Many cultivated for herbs, oils (Lavandula , lavender; Mentha , mint; Ocimum , basil; Rosmarinus , rosemary; Thymus , thyme), some cultivated as ornamental (in CA Cedronella , Leonotis , Phlomis )
Reference: [Cantino & Sanders 1986 Syst Bot 11:163–185]



Perennial from rhizomes, glabrous or hairy
Stems erect, branched or not
Leaves short-petioled to sessile, generally ovate to lanceolate; margin toothed to deeply lobed or cut below middle
Inflorescences axillary, each head-like, subtended by leaves
Flower: calyx generally 5-lobed, lobes ± equal, obtuse to short-awned; corolla slightly bilateral, not 2-lipped, generally 4-lobed, lobes ± unequal, odd lobe notched or entire; stamens 2, exserted, staminodes 2, minute, club-shaped; style exserted
Fruit: nutlets ± compressed, edge corky-thickened, truncate or rounded
Species in genus: 14 species: temp North America, Eurasia, 1 species in Australia
Etymology: (Greek: wolf foot, from French common name)
Reference: [Henderson 1962 Amer Midl Naturalist 68:95–135]


L. asper Greene

Rhizomes thicker and tuber-like near tip
Stems erect, 3–8(10) dm, puberulent to short-hairy
Leaf 2.5–7(9) cm, ± sessile, lanceolate to narrowly elliptic, serrate, glabrous to puberulent
Flower: calyx lobes awl-like, acuminate to short-awned; corolla 3–5 mm, slightly > calyx, white
Fruit: nutlet 1.5–2 mm; top truncate, sometimes minutely toothed
Chromosomes: 2n=22
Ecology: Uncommon. Moist areas, marshes, streambanks
Elevation: < 1300 m.
Bioregional distribution: Deltaic Great Central Valley, San Francisco Bay Area, Great Basin Floristic Province
Distribution outside California: to w Canada, Great Plains
Flowering time: Jun–Oct
Synonyms: L. lucidus Benth. misapplied

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