ERICACEAE

HEATH FAMILY

Gary D. Wallace, except as specified

Perennial, shrub, tree
Stem: bark often peeling distinctively
Leaves simple, generally cauline, alternate, opposite, rarely whorled, evergreen or deciduous, often leathery, petioled or not; stipules 0
Inflorescence: raceme, panicle, cyme, or flowers solitary, generally bracted; pedicels often with 2 bractlets
Flower generally bisexual, generally radial; sepals generally 4–5, generally free; petals generally 4–5, free or fused; stamens 8–10, free, filaments rarely appendaged, anthers awned or not, dehiscent by pores or slits; nectary generally at ovary base, disk-like; ovary superior or inferior, chambers generally 1–5, placentas axile or parietal, ovules 1–many per chamber, style 1, stigma head- to funnel-like or lobed
Fruit: capsule, drupe, berry
Seeds generally many, sometimes winged
Genera in family: ± 100 genera, 3000 species: generally worldwide except deserts; some cultivated, especially Arbutus, Arctostaphylos, Rhododendron, Vaccinium
Reference: [Wallace 1975 Wasmann J Biol 33:1–88; 1975 Bot Not 128:286–298]
Subfamilies Monotropoideae, Pyroloideae, Vaccinioideae sometimes treated as families. Nongreen plants obtain nutrition from green plants through fungal intermediates.

VACCINIUM

BLUEBERRY, HUCKLEBERRY, CRANBERRY

Shrub, tree, glabrous to hairy, rhizomed or not; burls generally 0
Stem trailing to erect
Leaves alternate
Inflorescence: raceme or flowers solitary, bracted; bud scales present; bractlets generally 2
Flower: sepals 4–5, 2/3 to fully fused (lobes then 0); petals generally 4–5, ± 2/3 fused, cylindric to urn- or cup-shaped, generally white; stamens 8 or 10, filaments generally glabrous, anthers elongate, dehiscent by pores on small tubes, awned or not; ovary inferior, chambers 4–5 (or appearing 10 by intrusion of ovary wall), placentas axile, stigma head-like
Fruit: berry
Seeds generally many
Species in genus: 400+ species: temp n hemisphere, tropical mtns, Africa
Etymology: (Latin: for V. myrtillus L.)
[Vander Kloet 1988 The genus Vaccinium in North America]

Introduced

V. macrocarpon Aiton

CRANBERRY

Shrub, glabrous to hairy, rhizomed
Stems prostrate and erect, < 1.5 dm, slender, rooting; twigs not angled, brownish
Leaves evergreen, generally 7–17 mm; blade narrowly elliptic to oblong, leathery, entire, lower surface glaucous
Inflorescence: flowers solitary in axils of reduced lowest leaves of youngest shoots; bractlets 2, leaf-like; pedicels recurved, jointed to flower
Flower: sepals 4, fused; petals 4, fused at base, lobes >> tube, reflexed when anthers open; filaments ± hairy, anthers unawned
Fruit 9–14 mm diam, red
Chromosomes: 2n=24
Ecology: Boggy soil at abandoned placer mine
Elevation: ± 900 m.
Bioregional distribution: n High Sierra Nevada (near North Columbia, n Nevada Co.)
Distribution outside California: native to e N.America