Jepson Flora Project: Jepson Interchange    

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Jepson Interchange (more information)
©Copyright 1993 by the Regents of the University of California

  • Up-to-date information about California vascular plants is available from the Jepson eFlora.



Dieter H. Wilken, except as specified

Annual to tree, sometimes monoecious or dioecious, glandular or with bead-like hairs that collapse with age, becoming scaly or powdery
Stem often fleshy
Leaves generally alternate, entire to lobed; veins generally pinnate
Inflorescence: raceme, spike, catkin-like, or spheric cluster, or flower 1; bracts 0–few
Flower: sepals 1–5, often 0 in pistillate flowers, free or fused, generally persistent in fruit; petals 0; stamens 0–5; ovary generally superior, chamber 1, ovule 1, styles 1–3
Fruit: generally utricle
Seed 1, vertical (fruit compressed side-to-side) or horizontal (fruit compressed top-to-bottom)
Genera in family: 100 genera, 1300 species: worldwide, especially deserts, saline or alkaline soils; some cultivated for food (Beta , beets, chard; Chenopodium , quinoa)
Recent taxonomic note: Recently treated in expanded Amaranthaceae



Dean Taylor and Dieter H. Wilken

Annual (generally monoecious) to shrub (generally dioecious), often scaly
Leaves generally alternate, generally entire; lower generally ± short-petioled; upper generally sessile, ± reduced
Staminate inflorescence: spike or spheric cluster; bracts 0
Pistillate inflorescence: clusters to spike- or panicle-like; bracts 2 per fruit, free to fused, generally compressed, generally sessile
Staminate flower: calyx lobes 3–5; stamens 3–5
Pistillate flower: calyx ± 0; ovary ovoid to spheric, style branches 2
Seed generally erect
Species in genus: ± 250 species: temp to subtropical worldwide
Etymology: (Latin: ancient name)
Generally in alkaline or saline soils; some weedy; some accumulate selenium.


A. triangularis Willd.


Annual 3–9 dm
Stems branched from base, ascending to ± erect, finely white-scaly, becoming glabrous
Leaves: blades 10–70 mm, lanceolate to triangular-hastate, green, glabrous to sparsely fine-scaly, entire to wavy-dentate, base truncate to rounded; upper petioled
Inflorescence generally panicle-like; branches spike-like, dense
Pistillate inflorescence: bracts in fruit 3–7 mm, fused at base, ovate to deltate, base truncate, short-tubercled, finely dentate
Seed 1.5–2.5 mm, black
Ecology: Wet places, marshes
Elevation: < 300 m.
Bioregional distribution: North Coast, s Outer North Coast Ranges, Great Central Valley, Central Coast, South Coast, Channel Islands
Distribution outside California: to e N.America, Eurasia
Synonyms: A. patula L. subsp. hastata (L.) H.M. Hall & Clem. misapplied

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