Jepson Flora Project: Jepson Interchange    

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Jepson Interchange (more information)
©Copyright 1993 by the Regents of the University of California

  • Up-to-date information about California vascular plants is available from the Jepson eFlora.



Alan R. Smith and Thomas Lemieux

Perennial, in soil or rock crevices; rhizome generally short-creeping, suberect, or erect, scales large, generally tan to brown, generally 1-colored
Leaves generally tufted, 5–200+ cm, generally ± alike; petiole generally firm, base generally darker, with 2–many vascular strands; blade 1–4-pinnate, often with scales, hair-like scales, hairs (except clear, needle-like hairs generally 0), or short-stalked glands on axes, sometimes between veins, veins free to netted; 1° and 2° axes generally grooved on upper side
Sporangia: sori round, less often oblong or J-shaped, along or at tips of veins; indusia peltate, round-reniform, oblong to linear, J-shaped, hood-like, or cup-like, rarely 0; spores elliptic, winged, ridged, or spiny, scar linear
Genera in family: ± 60 genera, > 1000 species: worldwide, especially tropical, wooded areas. Woodsia sometimes in Woodsiaceae; Athyrium, Cystopteris sometimes in Athyriaceae.



Rhizome generally suberect to erect, often stout
Leaf: petiole stout, firm, generally densely scaly, in X -section with many round vascular strands in an arc; blade generally 1–3-pinnate, thin to leathery, scaly, veins generally free, rarely casually joined; 1° leaflet bases often wider on distal side; teeth generally including bristle-like tips that are < 2 mm
Sporangia: sori round; indusium generally peltate, sinus 0
Species in genus: ± 175+ species: ± worldwide
Etymology: (Greek: many rows, from rows of sori on type sp.)


P. imbricans (D.C. Eaton) D.H. Wagner

Leaf: petiole generally 1/5–1/2 blade length, base scales ± 2–3 mm wide, lanceolate, those above lowest 1° leaflets generally < 1 mm wide, falling early; blade narrow-lanceolate to -elliptic, 1-pinnate
Chromosomes: 2n=82
Ecology: Shaded or exposed outcrops, banks, slopes, rocky areas
Elevation: 300–2500 m.
Bioregional distribution: Klamath Ranges, North Coast Ranges, High Cascade Range, n Sierra Nevada Foothills, High Sierra Nevada, San Francisco Bay Area, Outer South Coast Ranges, South Coast, Transverse Ranges, Peninsular Ranges, Modoc Plateau
Distribution outside California: to British Columbia
Hybridizes with P. dudleyi (called P. californicum ), P. lemmonii (called P. scopulinum ), P. munitum.

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bioregional map for POLYSTICHUM%20imbricans being generated

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