Grimmia lisae

   Plants to 1.5 cm high, in hoary green to gray-green cushions, opaque-green to nearly black below. Leaves loosely imbricate when dry, loosely erect-spreading when moist, somewhat concave but not at all keeled, to 3 mm long with the awn constituting nearly 1/2 of that length, ovate-lanceolate from a loosely sheathing somewhat expanded base, 2.5–3.5: 1. Median leaf cells in regular longitudinal rows, unistratose in the expanded base but uniformly bistratose in the limb, smooth, to 7 µm wide, isodiametric with lumens irregular but without corner thickenings. Marginal cells not differentiated. Basal juxtacostal cells elliptical, to 3: 1, to 15 µm wide, with irregularly thickened lateral walls. Basal marginal cells transversely elongate, 0.5–0.7: 1, to 15 µm wide. Costa flared at the base and filling 1/3–1/4 of immediate leaf base, narrowed and obscure above, percurrent in the somewhat acuminate awn. Costa cross-section at leaf middle broad and flattened, less than 1.5 times as thick as the adjacent leaf, with a weak abaxial band of stereids or substereids and with a single layer of larger cells on the adaxial face. Margin of leaf limb plane to incurved, entire below. Awn strongly decurrent with its margin somewht serrulate, and with its surface spinose-prorate. Axillary hairs to 100 µm long, with 6–8 nearly isodiametric and thick- walled cells, without a basal brown cell. Rhizoids red brown, to 15 µm wide at base, smooth. Stem cross-section rounded-triangular with a large and well-defined central strand, and with leptodermous, hyaline and large inner cortical cells, and with the 1–2 cell-thick outer stereome composed of much smaller, thick-walled cells strongly pigment with yellow to yellow-brown.

   Dioicous with male and female plants similar. Perichaetial leaves loosely sheathing base of seta, somewhat larger and with a relatively longer, less decurrent awn than the adjacent vegetative leaves. Seta to 2.5 mm long, yellow to pale brown, straight and smooth. Urn erect, symmetrical, smooth to lightly wrinkled when dry, to 1.2 mm long, 1.5–2: 1, not strumose. Operculum short rostrate, about 1/3 as long as urn. Annulus formed of 2–3 rows of relatively more thin-walled cells. Calyptra mitrate, covering about 1/2 of urn. Peristome red-brown, to 200 µm long, cribrose along median line near the apex, papillose to vertically striate-papillose. Exothecial cells to 25 µm wide, not in regular vertical rows, quite irregular in size and shape, rectangular 2–4: 1, rather thick walled. Stomata phaneroporous, restricted to base of urn. Spores smooth, to 12 µm.

Vouchers: Lake Co.: County Road to Lake Pillsbury at Dashiells Creek, Mendocino National Forest, Toren 7262 (CAS) [determined by Whittemore]; San Francisco Co.: Yerba Buena Island, Toren 8056 (CAS) and Shevock 19109 (determined by Muñoz); Santa Barbara Co.: Windmill Canyon, Santa Rosa Island, Channel Islands National Park, Shevock 20898 (determined by Muñoz); Santa Clara Co.: north shore of Lake Grant, Lake Grant County Park, Whittemore 6069 (CAS, MO) [determined by Muñoz]; Tulare Co.: along flume above Ash Mountain Headquarters, Sequoia National Park, Shevock & Morosco 16780 (determined by Muñoz).

Literature: Greven 2003; Kellman 2003; Shevock and Toren 2001; Whittemore and Sommers 1999. As Grimmia californica Brandegee 1891; Kingman 1912; Lesquereux 1868; Lesquereux and James 1884; Moxley 1928; Sullivant 1856; Watson 1880. As Grimmia trichophylla var. brachycarpa Toren 1977.

Illustration References: Greven 2003.

No illustrations found in database.

Elevation by latitude plot for Grimmia lisae
   in California