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Orthodicranum tauricum (Sapehin) Smirnova [Dicranaceae]


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Database links

UC Specimens in the University and Jepson Herbaria Public Portal

Specimens of Orthodicranum tauricum in the Consortium of North American Bryophyte Herbaria portal (CNABH)

Orthodicranum tauricum -- Dicranum tauricum Sapehin, Bot. Jahrb. 46, Beibl. 105: 10. 1911. -- Orthodicranum strictum (Mohr) Broth., Laubm. Fennosk. 92. 1923. -- Dicranum strictum Schleich. ex Mohr, Ann. Rot. 2: 546. 1806.

   Plants in light to dark green tufts or cushions to 4 cm high, rather glossy to almost lustreless, with the tips of most of the older leaves caducous. Leaves erect spreading wet or dry, not at all crispate when dry, secund or not, concave at the base and keeled to subtubular inrolled above, to 5 mm long, 8–12: 1, broadest immediately above the base, not plicate, lanceolate to linear lanceolate. Leaf lamina unistratose throughout or with bistratose streaks above, with median cells mostly short rectangular, to 12 µm wide, 1.2–4: 1, smooth, in regular rows, not pitted. Suprabasal cells, rectangular, to 8: 1, not well demarcated from the alar cells. Alar cells yellow brown, to 30 µm broad, pellucid but rather thick walled, at least in part quadrate. Costa filling about 1/5 of leaf at base, tapering and excurrent in the subulate apex. Costa cross-section with ad- and abaxial bands of substereids; with guide cells well demarcated but without well demarcated epidermises. Margin plane to ascending throughout, entire or with a few serrulations near apex. Rhizoids white to light brown, to 15 µm wide at base, smooth, sparingly branched, arising on stem from immediately below leaf insertion. Stem cross-section with central strand inconspicuous; with large and leptodermous interior cells and with only a slight differentiation of a partial layer of smaller pachydermous outer cortical cells.

   Dioicous with male and female plants similar. Perigonia terminal or both terminal and lateral in a periapical cluster. Perichaetial leaves erect to spreading, similar to adjacent vegetative leaves. Consistently monosetous with the seta to 25 mm, yellow brown, straight and smooth. Urn erect and symmetrical, cylindrical, to 2.5 mm long, about 3–4: 1, often irregularly wrinkled but neither sulcate nor strumose. Operculum rostrate, about 2/3 as long as urn. Annulus essentially undifferentiated. Peristome red-brown, inserted slightly below capsule mouth, to 300 µm long, divided below the middle into almost filiform segments, smooth to vertically low striate but occasionally papillose distally. Exothecial cells to 25 µm wide, in regular vertical rows, quite irregular in size and shape, rectangular 1–3: 1, rather thick walled with lumen/wall ratio about 2: 1. Spores to 15 µm, smooth to lightly papillose.




   This plant can be quite variable in both size and the degree of falcation of its leaves, In general, strongly falcate leaves can be found only on small plants growing on rotten logs. Its typical habitat is rotten logs and stumps but the largest plants (sometimes to 10 cm high are found on calcareous rocks in seepages at rather high elevations. These latter can have the leaves so strict that one is reminded of the more northern Dicranum elongatum Schleich. ex Schwaegr.


Vouchers: Del Norte Co.: between Wilderness Falls to Doe Flat, Norris 67761; El Dorado Co.: Sly Park Creek, Norris 58574; Marin Co.: Cataract Trail near Alpine Lake, Yurky 1050 (SFSU); Mendocino Co.: Highway 101 at Cummings turnoff, milepost 84.91, Norris 21639; Santa Cruz Co.: Zayante Canyon Road north of Felton, Norris 86874; Siskiyou Co.: Cedar Basin, South Fork Sacramento River, Shasta-Trinity National Forest, Shevock 12227; Tulare Co.: Redwood Mountain Grove, trail at Redwood Creek, Kings Canyon National Park, Shevock 17483.

Literature: Shevock and Toren 2001. As Dicranum strictum Howe 1896; Lesquereux 1868; Thomson and Ketchledge 1958; Watson 1880. As Dicranum tauricum Bourell 1981; Flowers 1973; Holmberg 1969; Jamieson 1969; Kellman 2003; Lawton 1971; McGrew 1976; Showers 1982; Smith 1970; Spjut 1971; Toren 1977; Yurky 1990. As Orthodicranum strictum Koch 1950a, 1951e; Koch and Ikenberry 1954.

Geographic subdivisions for
Orthodicranum tauricum: CaR, CW, NW, SN.
map of distribution

Illustration References: Malcolm et al. 2009 p. 109; Abramov and Volkova 1998; Enroth 1989; As Dicranum tauricum Flowers 1973; Ignatov and Ignatova 2003; Lawton 1971.

R. W. Spjut 1289

Orthodicranum tauricum
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© 2012 calbryos group
Orthodicranum tauricum
click for enlargement
© 2011 calbryos group

More photos of Orthodicranum tauricum in CalPhotos

Elevation by latitude plot for Orthodicranum tauricum
   in California