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Database links
UC Specimens in the University and Jepson Herbaria Public Portal
Specimens of Grimmia caespiticia in the Consortium of North American Bryophyte Herbaria portal (CNABH)
Grimmia caespiticia
Plants to 1.5 cm high, in hoary green to gray-green cushions, opaque-green to nearly black below. Leaves loosely imbricate when dry, loosely erect-spreading when moist, somewhat concave but not at all keeled, to 3 mm long with the awn constituting nearly 1/2 of that length, ovate-lanceolate from a loosely sheathing somewhat expanded base, 2.5–3.5: 1. Median leaf cells in regular longitudinal rows, unistratose in the expanded base but uniformly bistratose in the limb, smooth, to 7 µm wide, isodiametric with lumens irregular but without corner thickenings. Marginal cells not differentiated. Basal juxtacostal cells elliptical, to 3: 1, to 15 µm wide, with irregularly thickened lateral walls. Basal marginal cells transversely elongate, 0.5–0.7: 1, to 15 µm wide. Costa flared at the base and filling 1/3–1/4 of immediate leaf base, narrowed and obscure above, percurrent in the somewhat acuminate awn. Costa cross-section at leaf middle broad and flattened, less than 1.5 times as thick as the adjacent leaf, with a weak abaxial band of stereids or substereids and with a single layer of larger cells on the adaxial face. Margin of leaf limb plane to incurved, entire below. Awn strongly decurrent with its margin somewht serrulate, and with its surface spinose-prorate. Axillary hairs to 100 µmlong, with 6–8 nearly isodiametric and thick- walled cells, without a basal brown cell. Rhizoids red brown, to 15 µm wide at base, smooth. Stem cross-section rounded-triangular with a large and well-defined central strand, and with leptodermous, hyaline and large inner cortical cells, and with the 1–2 cell-thick outer stereome composed of much smaller, thick-walled cells strongly pigment with yellow to yellow-brown.
Dioicous with male and female plants similar. Perichaetial leaves loosely sheathing base of seta, somewhat larger and with a relatively longer, less decurrent awn than the adjacent vegetative leaves. Seta to 2.5 mm long, yellow to pale brown, straight and smooth. Urn erect, symmetrical, smooth to lightly wrinkled when dry, to 1.2 mm long, 1.5–2: 1, not strumose. Operculum short rostrate, about 1/3 as long as urn. Annulus formed of 2–3 rows of relatively more thin-walled cells. Calyptra mitrate, covering about 1/2 of urn. Peristome red-brown, to 200 µm. long, cribrose along median line near the apex, papillose to vertically striate-papillose. Exothecial cells to 25 µm. wide, not in regular vertical rows, quite irregular in size and shape, rectangular 2–4: 1, rather thick walled. Stomata phaneroporous, restricted to base of urn. Spores smooth, to 12 µm.
Vouchers: Inyo Co.: Big Pine Creek and Mt. Whitney Trail, Inyo National Forest, Greven s.n. (CAS); Mariposa Co.: Aqua Fria Road near Mariposa, Congdon s.n. (FH) [determined by Muñoz]; Mono Co.: Highway 182 at Devils Gate near Nevada State line,Toiyabe National Forest, Greven s.n. (CAS); Napa Co.: Napa Soda Springs, Mann s.n. (NY) [confirmed by Muñoz].
Literature: Muñoz 1998b. As. G. alpestris var. manniae Grout 1928-40.
Geographic subdivisions for Grimmia caespiticia: NW, SN, SNE. |
Illustration References: Greven 2003; Muñoz 1998b.
No illustrations found in database.
No expert verified images found for Grimmia caespiticia. |