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Amblystegium varium (Hedwig) Lindberg [Amblystegiaceae]


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Database links

UC Specimens in the University and Jepson Herbaria Public Portal

Specimens of Amblystegium varium in the Consortium of North American Bryophyte Herbaria portal (CNABH)

Amblystegium varium (Hedw.) Lindb., Musci Scand. 32. 1879 -- Leskea varia Hedwig, Spec. Musc. 216. 1801. -- Hygroamblystegium varium (Hedwig) Monk., Hedwigia 50: 275. Orthotheciella varia (Hedwig) Ochyra, Moss Flora of King George Island, Antarctica.

   Plants in loose and rather thin mats, pale to dull green. Stems to 3 cm long, prostrate to weakly ascending, irregularly branched. Stem leaves markedly larger and relatively more slender than branch leaves with all leaves erect-spreading to spreading when moist, little changed when dry. Leaves to 1.5 mm, 2.5–3: 1, symmetric and lanceolate to ovate-lanceolate, not at all plicate, long acuminate with the acumens straight. Leaf margins plane and entire or weakly serrulate in distal 1/3, not decurrent. Median cells smooth, to 8 µm wide, rhomboidal, 2–3: 1, rather thick-walled, not pitted. Alar cells isodiametric-rhomboidal with walls not strongly thickened, only gradually differentiated from adjacent laminal cells. Apical cells gradually more elongate than median cells. Costa extending into acumen, often somewhat sinuose distally, without an abaxial spine. Pseudoparaphyllia deltoid, strongly overarching the bud, occasionally singly placed and resembling paraphllia. Axillary hairs of 3 similar cells, to 50 µm with no basal brown cells, not offset from leaf insertion. Rhizoids light brown, to 16 µm, smooth, sparingly present on older prostrate stems.

   Autoicous with perigonia near perichaetia. Perichaetial bracts costate, lanceolate to narrowly lanceolate, about 4: 1 with cells uniformly long and straight to 10: 1. Seta brown to red-brown, smooth, to 2 cm long. Capsule brown to red brown, inclined, not strangulate. Urn to 1.8 mm, short cylindric to oblong, strongly arcuate, 3.5–4.5: 1. Exothecial cells in regular vertical rows, to 20 µm wide, 2: 1, mostly with oblique transverse walls, with moderately thick, red-brown walls and with stomata not seen in our specimens. Operculum conic to apiculate, about 1/5 as long as urn. Annulus differentiated in 1–2 rows of enlarged cells, not revoluble. Exostome to 500 µm long, horizontally lightly striate below, smooth above, yellow-brown. Endostome segments smooth and nodose, keel, mostly shorter than exostome with basal membrane about equal in length to the segments. Cilia 1–3, nodose. Calyptra cucullate, smooth, without hairs. Spores finely papillose, to 15 µm.






Vouchers: Kern Co.: Kern Plateau, Caldwell Creek near Kernville, Sequoia National Forest, Laeger 310 (CAS) and BLM Short Canyon ACEC west of Highway 14, Laeger & Davis 1290 (CAS) [determined by Hedenäs]; Mariposa Co.: in stream below Highway 49 about 3.3 miles southeast of Mariposa, Shevock 15238; San Bernardino Co.: East Fork Mountain Home Creek, San Bernardino Mountains, San Bernardino National Forest, Wheeler 8231 (CAS, UC); Santa Cruz Co.: along San Lorenzo River, Ben Lomond, Kellman 1477 (CAS); Tulare Co.: South Creek at South Creek Falls, Sequoia National Forest, Norris 87081.

Literature: Flowers 1973; Harthill et al. 1979; Kingman 1912; Koch 1950a; Mishler 1978; Toren 1977. As Orthotheciella varia Kellman 2003; Ochyra 1998a.

Geographic subdivisions for
Amblystegium varium: CW, DMoj, SN, SW.
map of distribution

Illustration References: Flowers 1973; Hedenäs 2003; Ireland 1982; Sharp et al. 1994.

No illustrations found in database.

No expert verified images found for Amblystegium varium.

Elevation by latitude plot for Amblystegium varium
   in California