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BRENT DRENNEN MISHLER


Director, University and Jepson Herbaria
Professor, Department of Integrative Biology
University of California, Berkeley

1001 Valley Life Sciences Building #2465
University of California
Berkeley, CA 94720-2465
phone: (510) 642-6810
FAX: (510) 643-5390
E-mail: bmishler@berkeley.edu
WWW: http://ucjeps.herb.berkeley.edu/people/mishler.html

DEGREES:

PROFESSIONAL EMPLOYMENT:

GRANTS AND AWARDS:

RESEARCH INTERESTS:

My research interests can be grouped into two main areas: empirical studies of ecology, phylogeny, systematics, and development of mosses, and the theoretical basis of systematic and evolutionary biology. Empirical studies include: (1) the phylogenetic relationships of the major groups of bryophytes (including mosses) and other land plants, using morphological, developmental, and ultrastructural characters as well as chloroplast DNA sequence data; (2) the development of moss peristomes in relation to evolution of the group; and (3) biosystematic studies of the haplolepideous mosses, including the tropical family Calymperaceae and the diverse temperate genus Tortula (Syntrichia), which involve transplant and ecological studies in the field, comparative physiological measurements and culture experiments in the lab, and morphological studies in the herbarium; (4) the reproductive biology of bryophytes, especially dryland mosses; and (5) the bryophyte flora of California and of Moorea (in the Society Islands of the South Pacific). Theoretical studies include investigations of the nature of species and speciation, methods for phylogenetic reconstruction (with an emphasis on cladistic analysis of molecular data), and the relationship between development and evolution.

BIBLIOGRAPHY:

Peer-Reviewed Articles in Professional Journals and Books:

1979.
M.P. Harthill, D.M. Long and B.D. MISHLER. Preliminary list of southern California mosses. The Bryologist 82: 260-267.
1982.
B.D. MISHLER and M.J. Donoghue. Species concepts: a case for pluralism. Systematic Zoology 31: 491-503. **[Reprinted 1992 in: Ereshefsky, M. (ed.) The Units of Evolution: Essays on the Nature of Species. MIT Press.] **[Reprinted 1994 in: Sober, E. (ed.) Conceptual Issues in Evolutionary Biology (second edition). MIT Press.]
1983
B.D. MISHLER and N.G. Miller. Distributional studies of Massachusetts bryophytes. Rhodora 85: 421-432.
1984
B.D. MISHLER and S.P. Churchill. A cladistic approach to the phylogeny of the "bryophytes". Brittonia 36: 406-424.
1985
B.D. MISHLER and S.P. Churchill. Cladistics and the land plants: a response to Robinson. Brittonia 37: 282-285.

B.D. MISHLER. Biosystematic studies of the Tortula ruralis complex. I. Variation of taxonomic characters in culture. Journal of the Hattori Botanical Laboratory 58: 225-253.

B.D. MISHLER. The morphological, developmental, and phylogenetic basis of species concepts in bryophytes. The Bryologist 88: 207-214.

B.D. MISHLER and S.P. Churchill. Transition to a land flora: phylogenetic relationships of the green algae and bryophytes. Cladistics 1: 305-328.

B.D. MISHLER. The phylogenetic relationships of Tortula: an SEM survey and a preliminary cladistic analysis. The Bryologist 88: 388-403.

1986
B.D. MISHLER. Ontogeny and phylogeny in Tortula (Musci: Pottiaceae). Systematic Botany 11: 189-208.

B.D. MISHLER. A Hennigian approach to bryophyte phylogeny. Journal of Bryology 14: 71-81.

1987
B.D. MISHLER. Sociology of science and the future of Hennigian phylogenetic systematics. Cladistics 3: 55-60.

B.D. MISHLER and S.P. Churchill. Transition to a land flora: a reply. Cladistics 3: 65-71.

K. Bremer, C.J. Humphries, B.D. MISHLER, and S.P. Churchill. On cladistic relationships in green plants. Taxon 36: 339-349.

B.D. MISHLER. Leaf development in Tortula papillosissima (Pottiaceae). Mem. New York Bot. Gard. 45: 48-54.

J. Shaw, L.E. Anderson, and B.D. MISHLER. Peristome development in mosses in relation to systematics and evolution. I. Diphyscium foliosum (Buxbaumiaceae). Mem. New York Bot. Gard. 45: 55-70.

B.D. MISHLER and R.N. Brandon. Individuality, pluralism, and the phylogenetic species concept. Biology and Philosophy 2: 397-414. **[Reprinted 1996 in: Brandon, R.N. Concepts and Methods in Evolutionary Biology . Cambridge University Press.] 1998 in: Hull, D.L. and Ruse, M. The Philosphy of Biology. Oxford University Press.]

1988
B.D. MISHLER. Relationships between ontogeny and phylogeny, with reference to bryophytes. In C. J. Humphries (ed.), Ontogeny and Systematics, pp. 117-136. Columbia University Press.

B.D. MISHLER. Reproductive ecology of bryophytes. In J. Lovett Doust and L. Lovett Doust (eds.), Plant Reproductive Ecology, pp. 285-306. Oxford University Press.

B.D. MISHLER, K. Bremer, C.J. Humphries, and S.P. Churchill. The use of nucleic acid sequence data in phylogenetic reconstruction. Taxon 37: 391-395.

B.D. MISHLER and A.E. Newton. Influences of mature plants and desiccation on germination of spores and gametophytic fragments of Tortula. Journal of Bryology 15: 327-342.

1989
J. Shaw, L.E. Anderson, and B.D. MISHLER. Peristome development in mosses in relation to systematics and evolution. III. Funaria hygrometrica, Bryum pseudocapillare, and B. bicolor. Systematic Botany 14:24-36.

B.D. MISHLER and R.N. Brandon. Sex and the individuality of species: a response to Ghiselin. Biology and Philosophy 4:77-79.

J. Shaw, B.D. MISHLER, and L.E. Anderson. Peristome development in mosses in relation to systematics and evolution. IV. Haplolepideae: Ditrichaceae and Dicranaceae. The Bryologist 92: 314-325.

1990
B.D. MISHLER and A.F. Budd. Species and evolution in clonal organisms-- introduction. Systematic Botany 15: 79-85.

B.D. MISHLER. Reproductive biology and species distinctions in the moss genus Tortula, as represented in Mexico. Systematic Botany 15: 86-97.

A.F. Budd and B.D. MISHLER. Species and evolution in clonal organisms--summary and discussion. Systematic Botany 15: 166-171.

1991
B.D. MISHLER and M.J. Oliver. Gametophytic phenology of Tortula ruralis, a desiccation-tolerant moss, in the Organ Mountains of southern New Mexico. The Bryologist 94: 143-153.

B.D. MISHLER. Phylogenetic analogies in the conceptual development of science. PSA 1990, Volume Two, pp. 225-235. [Proceedings of the 1990 biennial meeting of the Philosophy of Science Association]

B.D. MISHLER and E. De Luna. The use of ontogenetic data in phylogenetic analyses of mosses. Advances in Bryology 4: 121-167.

L.E. Graham, C.F. Delwiche, and B.D. MISHLER. Phylogenetic connections between the "green algae" and the "bryophytes". Advances in Bryology 4: 213-244.

1992
V.A. Albert, B.D. MISHLER, and M.W. Chase. Character-state weighting for restriction site data in phylogenetic reconstruction, with an example from chloroplast DNA. In P. Soltis, D. Soltis, and J. Doyle (eds.), Molecular Systematics of Plants, pp. 369-403. Chapman and Hall.

B.D. MISHLER, P.H. Thrall, J.S. Hopple, Jr., E. De Luna, and R.J. Vilgalys. A molecular approach to the phylogeny of bryophytes: cladistic analysis of chloroplast-encoded 16S and 23S ribosomal RNA genes. The Bryologist 95: 172- 180.

V.A. Albert and B.D. MISHLER. On the rationale and utility of weighting nucleotide sequence data. Cladistics 8: 73-83.

1993
M.J. Oliver, B.D. MISHLER, and J.E. Quisenberry. Comparative measures of desiccation-tolerance in the Tortula ruralis complex. I. Variation in damage control and repair. American Journal of Botany 80: 127-136.

M.W. Chase, D.E. Soltis, R.G. Olmstead, D. Morgan, D.H. Les, B.D. MISHLER, M.R. Duvall, R.A. Price, H.G. Hills, Y. Qiu, K.A. Kron, J.H. Rettig, E. Conti, J.D. Palmer, J.R. Manhart, K.J. Sytsma, H.J. Michaels, W.J. Kress, K.G. Karol, W.D. Clark, M. Hedrén, B.S. Gaut, R.K. Jansen, K. Kim, C.F. Wimpee, J.F. Smith, G.R. Furnier, S.H. Strauss, Q. Xiang, G.M. Plunkett, P.S. Soltis, S.M. Swensen, S.E. Williams, P.A. Gadek, C.J. Quinn, L.E. Eguiarte, E. Golenberg, G.H. Learn, Jr., S.W. Graham, S.C.H. Barrett, S. Dayanandan, and V.A. Albert. Phylogenetics of seed plants: an analysis of nucleotide sequences from the plastid gene rbcL. Annals Missouri Botanical Garden 80: 528-580.

V.A. Albert, M.W. Chase, and B.D. MISHLER. Character-state weighting for cladistic analysis of protein-coding DNA sequences. Annals Missouri Botanical Garden 80: 752-766.

1994
B.D. MISHLER. Tortula. In A.J. Sharp, H. Crum, and P. M. Eckel (eds.), The Moss Flora of Mexico. Mem. New York Bot. Gard. 69: 319-352. [a monographic treatment of 25 species]

B.D. MISHLER. Cladistic analysis of molecular and morphological data. American Journal of Physical Anthropology 94: 143-156.

B.D. MISHLER, L.A. Lewis, M.A. Buchheim, K.S. Renzaglia, D.J. Garbary, C.F. Delwiche, F.W. Zechman, T.S. Kantz, and R.L. Chapman. Phylogenetic relationships of the "green algae" and "bryophytes." Annals Missouri Botanical Garden 81: 451-483.

V.A. Albert, A. Backlund, K. Bremer, M.W. Chase, J.R. Manhart, B.D. MISHLER, and K.C. Nixon. Functional constraints and rbcL evidence for land plant phylogeny. Annals Missouri Botanical Garden 81: 534-567.

R.N. Brandon, J. Antonovics, R. Burian, S. Carson, G. Cooper, P.S. Davies, C. Horvath, B.D. MISHLER, R.C. Richardson, K. Smith, and P. Thrall. Discussion: Sober on Brandon on screening-off and the levels of selection. Philosophy of Science 61: 475-486.

A.E. Newton and B.D. MISHLER. The evolutionary significance of asexual reproduction in mosses. Journal of the Hattori Botanical Laboratory 76: 127-145.

1995
B.D. MISHLER, B.G. Baldwin, and S. D'Alcamo. The future of California floristics and systematics: research, education, conservation. Madroño 42: 93-95.

B.D. MISHLER. Plant systematics and conservation: science and society. Madroño 42: 103-113.

1996
B.D. MISHLER, V. A. Albert, M. W. Chase, P. O. Karis, and K. Bremer. Character- state weighting for DNA restriction site data: asymmetry, ancestors, and the Asteraceae. Cladistics 12: 11-19.

E. De Luna and B.D. MISHLER. El concepto de homología filogenética y la selección de caracteres taxonómicos. Boletín de la Sociedad Botánica de México 59: 131-146.

1997
L.A. Lewis, B.D. MISHLER, and R. Vilgalys. Phylogenetic relationships of the liverworts (Hepaticae), a basal embryophyte lineage, inferred from nucleotide sequence data of the chloroplast gene rbcL. Molecular Phylogenetics and Evolution 7: 377-393.

B.D. MISHLER and E. De Luna. Sistemática filogenética y el concepto de especie. Boletín de la Sociedad Botánica de México 60: 45-57.

1998
L. R. Stark, B.D. MISHLER, and D.N. McLetchie. Sex expression and growth rates in natural populations of the desert soil crustal moss Syntrichia caninervis. Journal of Arid Environments 40: 401-416.
1999
B.D. MISHLER. Getting rid of species? In R. Wilson (ed.), Species: New Interdisciplinary Essays, pp.307-315. MIT Press.

B. Goffinet, J. Shaw, L.E. Anderson, and B.D. MISHLER. Peristome development in mosses in relation to systematics and evolution. V. Diplolepideae: Orthotrichaceae. The Bryologist 102: 581-594.

E. De Luna, A.E. Newton, A. Withey, D. Gonzalez, and B.D. MISHLER. The transition to pleurocarpy: a phylogenetic analysis of the main Diplolepideous lineages based on rbcL sequences and morphology. The Bryologist 102: 634-650.

2000.
J. Shaw, L.E. Anderson, and B.D. MISHLER. Paedomorphic sporophyte development in Bruchia flexuosa (Bruchiaceae). The Bryologist 103: 147-155.

M.A. Bowker, L. R. Stark, D.N. McLetchie, and B.D. MISHLER. Sex expression, skewed sex ratios, and microhabitat distribution in the dioecious desert moss Syntrichia caninervis (Pottiaceae). American Journal of Botany 87: 517-526.

A.E. Newton, C.J. Cox, J.G. Duckett, J. Wheeler, B. Goffinet, T.A.J. Hedderson, and B.D. MISHLER. Evolution of the major moss lineages: phylogenetic analyses based on multiple gene sequences and morphology. The Bryologist 103: 187- 211.

C. La Farge, B.D. MISHLER, J. Wheeler, D. Wall, K. Johannes, S. Schaffer, and J. Shaw. Phylogenetic relationships within the haplolepideous mosses. The Bryologist 103: 257-276.

B.D. MISHLER and E. Theriot. The phylogenetic species concept sensu Mishler and Theriot: monophyly, apomorphy, and phylogenetic species concepts. In Q.D. Wheeler & R. Meier (eds.), Species Concepts and Phylogenetic Theory: A Debate, pp.44-54. Columbia University Press.

B.D. MISHLER and E. Theriot. A critique from the Mishler and Theriot phylogenetic species concept perspective: monophyly, apomorphy, and phylogenetic species concepts. In Q.D. Wheeler & R. Meier (eds.), Species Concepts and Phylogenetic Theory: A Debate, pp.119-132. Columbia University Press.

B.D. MISHLER and E. Theriot. A defense of the phylogenetic species concept sensu Mishler and Theriot: monophyly, apomorphy, and phylogenetic species concepts. In Q.D. Wheeler & R. Meier (eds.), Species Concepts and Phylogenetic Theory: A Debate, pp.179-184. Columbia University Press.

In Press:

B.D. MISHLER. Biodiversity and the loss of lineages. In L. Maffi (ed.), On Biocultural Diversity: Linking Language, Knowledge, and the Environment. Smithsonian Institution Press.

L. R. Stark, B.D. MISHLER, and D.N. McLetchie. The cost of realized sexual reproduction: assessing patterns of reproductive allocation and sporophyte abortion in a desert moss. American Journal of Botany

M.J. Oliver, Z. Tuba, and B.D. MISHLER. The evolution of vegetative desiccation tolerance in land plants. Plant Ecology

J. A. Wheeler, D. P. Wall, K. Johannes, and B.D. MISHLER. Congruence and convergence in the moss family Calymperaceae: phylogenetic analysis of two chloroplast genes (rbcL and rps4) and morphology. Systematic Biology.

I. Granzow-De La Cerda and B.D. MISHLER. A phylogenetic reconsideration of the circumscription of Desmatodon Brid. (Musci: Pottiaceae) in North America and Europe. The Bryologist

Supervised Papers:

[Note that it is my policy not to co-author papers resulting from my graduate students' dissertations, because I encourage them to work independently. Following are dissertation papers that I have supervised and provided funding for as major professor]

De Luna, E. 1990. Protonemal development in the Hedwigiaceae (Musci), and its systematic significance. Systematic Botany 15: 192-204.

De Luna, E. 1990. Developmental evidence of acrocarpy in Hedwigia ciliata (Musci: Hedwigiaceae). Tropical Bryology 2: 53-60.

De Luna, E. 1995. The circumscription and phylogenetic relationships of the Hedwigiaceae (Musci). Systematic Botany 20: 347-373.

Murrell, Z.E. 1993. Phylogenetic relationships in Cornus (Cornaceae). Systematic Botany 18: 469-495.

Murrell, Z.E. 1994. Dwarf dogwoods: intermediacy and the morphological landscape. Systematic Botany 19: 539-556.

Murrell, Z.E. 1996. A new section of Cornus in South and Central America. Systematic Botany 21: 273-288.

Newton, A.E. In Press. Three new species, and nomenclatorial notes and changes in Pireella (Pterobryaceae: Musci). Novon.

Newton, A.E. In Press. Monographic treatment of the genus Pireella (Pterobryaceae: Musci). Flora Neotropica Monographs.

Pryer, K.M., A.R. Smith, and J.E. Skog. 1995. Phylogenetic relationships of extant ferns based on evidence from morphology and rbcL sequences. American Fern Journal 85: 205-282.

Pryer, K.M. 1999. Phylogeny of Marsileaceous ferns and relationships of the fossil Hydropteris pinnata reconsidered. International Journal of Plant Sciences 160: 931-954.

Rice, S.K. and P.H. Schuepp. 1995. On the ecological and evolutionary significance of branch and leaf morphology in aquatic Sphagnum (Sphagnaceae). American Journal of Botany 82: 833-846.

Rice, S.K. 1995. Patterns of allocation and growth in aquatic Sphagnum species. Canadian Journal of Botany 73: 349-359.

Rice, S.K. and L. Giles 1996. The effects of leaf anatomy and water content on carbon isotope discrimination and photosynthesis in Sphagnum. Plant, Cell and Environment 19: 118-124.

Schwartz, O.M. 1991. A history of developmental studies of peristomes. Evansia 8: 55-73.

Schwartz, O.M. 1994. The development of the peristome-forming layers in the Funariaceae. International Journal of Plant Sciences 155: 640-657.

Schwartz, O.M. 1997. Apical cell segmentation and its relationship to the peristome-forming layers in the Funariaceae. International Journal of Plant Sciences 158:236-248.

Withey, A. 1996. Phylogenetic studies of the Spiridentaceae (Musci): observations of three morphological characters associated with pleurocarpy. Anales Instituto de Biología, UNAM, ser Botánica 67: 5-14.

Book Reviews and Popular Articles (non-peer reviewed):

1983.B.D. MISHLER. Book review: an illustrated moss flora of the maritimes. Rhodora 85: 391-394.

1984. B.D. MISHLER. Bryophyte ecology (book review). American Scientist 72: 201.

1985. B.D. MISHLER. Review: Thomas Duncan and Tod F. Stuessy (Editors). Cladistics: perspectives on the reconstruction of evolutionary history. The Bryologist 88: 420- 421.

B.D. MISHLER. Bryophytes of Northeastern North America. Newsletter of the Friends of the Farlow 3(2): 3-4.

1988. L.E. Anderson and B.D. MISHLER. Bryology at Duke University. Bryological Times 46: 1-3.

B.D. MISHLER. Species. Newsletter of the Friends of the Farlow 13: 1-3.

1989. B. Chernoff, T. Duncan, T. Elias, B.D. MISHLER, M. Novacek, E. Theriot, Q. Wheeler, and D.S. Wood. Commentary on the place of systematics. Association of Systematics Collections Newsletter 17(1):1-2.

B.D. MISHLER. Review: Science as a process: an evolutionary account of the social and conceptual development of science, by David L. Hull. Systematic Botany 14: 266-268.

1990. B.D. MISHLER. Review: R.N. Chopra and P.K. Kumra. Biology of bryophytes. The Bryologist 93: 100.

B.D. MISHLER. Species, speciation, and phylogenetic systematics. Cladistics 6: 205- 209.

1992. B.D. MISHLER, L.E. Anderson, and M. McMullen. Bryology at Duke University (with complete bibliography). Cryptogamic Herbarium, Duke University.

1993. B.D. MISHLER. Plant systematics and the future of the Jepson Herbarium: a view from the new director. The Jepson Globe 5(1): 1-5.

1994. B.D. MISHLER. Phylogeny and its importance for practical botany and conservation. The Jepson Globe 5(3) : 1-3.

1995. B.D. MISHLER. New connections for the Jepson Herbarium. The Jepson Globe 6(1) : 1-4.

B.D. MISHLER. The Molecular Phylogenetics Laboratory. The Jepson Globe 6(3) : 1-3.

1996. B.D. MISHLER. Spring field trips of the Jepson Herbarium. The Jepson Globe 7(2) : 1-4.

1997. B.D. MISHLER. Current events at the Jepson Herbarium: Bacigalupi gift, PEET Grant, and the Jepson Flora Project. The Jepson Globe 8(1) : 1-7.

1998. B.D. MISHLER. An external review of University and Jepson Herbaria. The Jepson Globe 9(2): 1-3.

B.D. MISHLER, H. Mohamed, and B. Tan. PEET mossing in Southeast Asia: collaborative fieldwork in Malaysia and Singapore. Bryological Times 97: 1-2.

1999. J.H. Lipps, C. D'Antonio, K. Grove, C. Hickman, B.D. MISHLER, and Z Wang. Geology and natural history of the Central Coast Ranges: Berkeley to Bodega Head, California. California Division of Mines and Geology Special Publication 119: 188-201.

B.D. MISHLER. New staff -- new directions for the Friends. The Jepson Globe 10(1): 1-2.

B.D. MISHLER. Bryophyte and fern studies in the Bryolab at the University and Jepson Herbaria. The Jepson Globe 10(3): 1-5.

Presented Papers and Symposia (most with published abstracts):

1977. A preliminary study on the effect of air pollution on the distribution of Orthotrichum in southern California. Annual meeting, American Bryological and Lichenological Society (ABLS). East Lansing, Michigan.

Biosystematics as applied to mosses. Annual meeting, Southern California Academy of Sciences. Pomona, California.

1979. Flora of the San Gabriel Mountains, California: Bryophytes. Annual meeting, ABLS. Stillwater, Oklahoma. (With David M. Long.)

1981. Studies on Tortula amphidiacea (C. Muell.) Broth. and T. fragilis Tayl., two Appalachian mosses also found in Middle America. Annual meeting, ABLS. Highlands, North Carolina.

1982. Species concepts and phylogenetic systematics. Annual meeting, Willi Hennig Society. College Park, Maryland.

1983. Phylogenetic relationships and generic limits of Tortula (Musci: Pottiaceae): evidence from SEM studies of the peristome. Annual meeting, American Society of Plant Taxonomists (ASPT). Grand Forks, North Dakota.

Development of the leaf of Tortula obtusissima: systematic and ecological implications. Annual meeting, ABLS. Grand Forks, North Dakota. (With Daniel C. Scheirer.)

On the application of Hennigian cladistics to the phylogeny of the bryophytes. Annual meeting, ABLS. Grand Forks, North Dakota. (With Steven P. Churchill.)

1984. The morphological, developmental, and phylogenetic basis of species concepts in bryophytes. Annual meeting, ABLS. Fort Collins, Colorado. (An invited paper.)

Transition to a land flora: phylogenetic relationships and classification of the "green algae" and the "bryophytes". Annual meeting, Willi Hennig Society. London, England. (With Steven P. Churchill -- an invited paper.)

1985. Relationships between ontogeny and phylogeny in the mosses. Third International Congress of Systematic and Evolutionary Biology. Brighton, England. (An invited paper.)

A Hennigian approach to bryophyte phylogeny. Third International Congress of Systematic and Evolutionary Biology. Brighton, England. (An invited paper.)

Sociology of science and the future of Hennigian phylogenetic systematics. Annual meeting, Willi Hennig Society. Miami, Florida. (An invited pre-dinner address.)

1986. Biosystematic studies of the Tortula ruralis complex. Annual meeting, ABLS. Amherst, Massachusetts.

Gametophyte development in the Mniaceae (Musci) and its phylogenetic implications. Annual meeting, Willi Hennig Society. New York, NY. (With Steven P. Churchill.)

1987. Individuality, pluralism, and the phylogenetic species concept. Summer Conference on History, Philosophy, and Social Studies of Biology. Blacksburg, VA. (With R.N. Brandon.)

The application of cladistics to species and to higher-level taxa of Bryophytes. XIV International Botanical Congress. Berlin, Germany. (An invited paper.)

Germination of spores and gametophytic fragments of Tortula under a variety of experimental conditions. Annual meeting, ABLS. Columbus, Ohio. (With A.E. Newton)

1988. The role of cladistics in systematics and evolutionary theory. Annual meeting, Phycological Society of America. Asilomar, California. (An invited paper.)

Evolution of desiccation-tolerance in the Tortula ruralis complex. I. Distribution, habitats, and water relationships. II. Physiological and biochemical mechanisms. Annual meeting, ABLS. Davis, California. (With M.J. Oliver.)

El uso de informacion ontogenetica en la sistematica de las briofitas. I Simposio Latinamericano De Briologia. Bogota, Colombia. (With Efraîn De Luna.)

Phylogenetic relationships of Agaricales. Annual meeting, Willi Hennig Society.

Stockholm, Sweden. (With Rytas Vilgalys and David Hibbett--an invited paper.)

The nature of sexual and asexual species in the moss genus Tortula. Annual meeting, Society of Systematic Zoology. San Francisco, California. (An invited paper.)

1989. Systematics, ecology, and biogeography of Tortula in Mexico. Tropical Bryology Conference, International Association of Bryologists. St. Louis, Missouri. (An invited paper.)

Peristome development in mosses in relation to systematics and evolution. Annual meeting, ASPT. Toronto, Canada. (With J. Shaw and L.E. Anderson.)

Phylogenetic relationships in the "charophyte" green algae. Annual meeting, Willi Hennig Society. Ithaca, New York. (An invited paper--with C. Delwiche and L.E. Graham).

1990. Problems and prospects for phylogenetic reconstruction of Angiosperm relationships from rbcL data. Annual meeting, ASPT. Richmond, Virginia. (An invited paper--with V. A. Albert and M. W. Chase.)

Interspecific differences in the control and repair of desiccation-damage within the Tortula ruralis complex. Annual meeting, ABLS. Richmond, Virginia. (With M. J. Oliver.)

The emperor's new clothes? A molecular approach to the phylogeny of bryophytes using chloroplast DNA. Annual meeting, ASPT. Richmond, Virginia. (With P. Thrall, J. S. Hopple, Jr., E. De Luna, and R. Vilgalys.)

Integrating molecular and morphological data in phylogenetic reconstruction of the bryophytes. Annual meeting, ABLS. Wakulla Springs, Florida.

Invited participant in a symposium entitled "Science as a Process". Biennial meeting, Philosophy of Science Association. Minneapolis, Minnesota.

1991. Capsule developmental anatomy and the systematic position of the Hedwigiaceae (Musci). Annual meeting, ABLS. San Antonio, Texas. (With E. DeLuna.)

Uses of developmental information in phylogenetic reconstruction. Annual meeting, Botanical Society of America. San Antonio, Texas. (An invited paper.)

Maximum likelihood character-state weighting for parsimony analysis of nucleotide data: examples using characters from the plastid-encoded rbcL gene. Annual meeting, ASPT. San Antonio, Texas. (With V.A. Albert and M.W. Chase.)

The relevance of rbcL sequence data to the phylogeny of bryophytes. Annual meeting, ASPT. San Antonio, Texas. (With C. Colacino and M.W. Chase.)

Homology and the recognition of character states. Annual meeting, ASPT. San Antonio, Texas. (With E. DeLuna.)

An overview of competing concepts of species and speciation. Annual meeting, Mycological Society of America. San Antonio, Texas. (An invited paper.)

Phylogenetic classification in biology: A revolution in evolution. Biennial meeting, International Society for the History, Philosophy, and Social Studies of Biology (ISHPSSB). Evanston, Illinois.

Cladistics, morphogenetic constraints, and species concepts in plants. Biennial meeting, ISHPSSB. Evanston, Illinois.

The decay index as a measure of relative robustness within a cladogram. Annual meeting, Willi Hennig Society. Toronto, Canada. (With M.J. Donoghue and V.A. Albert.)

Monophyly: divergence, reticulation, and species. Annual meeting, Willi Hennig Society. Toronto, Canada. (An invited paper.)

Phylogenetic relationships within the bryophyta: a preliminary analysis based on rbcL data. 86th Congress of the Societa Botanica Italiana. Viterbo, Italy. (With C. Colacino and M.W. Chase.)

1992. The cladistic analysis of molecular and morphological data. Annual meeting, American Association of Physical Anthropologists. Las Vegas, Nevada. (An invited paper.)

Phylogenetic reconstruction in bryophytes using nucleotide sequence data from chloroplast DNA. Annual meeting, ABLS. Honolulu, Hawaii. (An invited paper.)

The application of nucleotide sequence data from the rbcL gene to the phylogeny of bryophytes. Annual meeting, ASPT. Honolulu, Hawaii. (With L.A. Lewis, D. Gonzalez, and R. Vilgalys.)

Phylogenetic relationships of the "green algae" and "bryophytes" to the tracheophytes. 39th Annual Systematics Symposium, Missouri Botanical Garden, St. Louis. (An invited paper--with L.A. Lewis, M.A. Buchheim, and R.L. Chapman.)

1993. Towards a unified phylogenetic species concept. Annual meeting, ASPT. Ames, Iowa. (An invited paper.)

Phylogenetic analysis of the liverworts (Bryophyta: Hepaticae) using morphological and molecular characters. Annual meeting, ASPT. Ames, Iowa. (With L.A. Lewis and R. Vilgalys.)

Compartmentalization: local versus global parsimony. Annual meeting, Willi Hennig Society. Fullerton, California.

Evolutionary roles of asexual reproduction in bryophytes. XV International Botanical Congress. Yokohama, Japan. (An invited paper--with A.E. Newton.)

Molecular systematics of bryophytes. XV International Botanical Congress. Yokohama, Japan. (An invited paper.)

Problems and prospects for the inference of land-plant relationships from rbcL sequence variation. XV International Botanical Congress. Yokohama, Japan. (An invited paper--with M.W. Chase, V.A. Albert, A. Backlund, K. Bremer, J.R. Manhart, and K.C. Nixon.)

1994. Phylogeny and ontogeny: heterochrony, ecology, and reproductive biology of mosses. 10th Annual Southwestern Botanical Systematics Symposium, Rancho Santa Ana Botanic Garden. Claremont, California. (An invited paper.)

The future of plant systematics, science, and society. Symposium: The Future of California Floristics and Systematics, Friends of the Jepson Herbarium. Berkeley, California. (An invited paper.)

The future of bryology in North America. Annual meeting, ABLS. Knoxville, Tennessee. (An invited paper.)

Speciation, adaptation, and evolution. 41st Annual Systematics Symposium, Missouri Botanical Garden, St. Louis. (An invited paper.)

1995. Integrating molecular data into systematics. Annual meeting, ABLS. Jasper National Park, Canada. (An invited paper.)

Introduction to bryophyte conservation issues. Center for Plant Conservation, California Plant Conservation Task Force Meeting. Berkeley, California. (With D.H. Norris.)

A phylogenetic analysis of the subclass Mezgeriidae Bartholomew-Began (Jungermanniopsida). International Association of Bryologists meeting. Mexico City, Mexico. (With B. Crandall-Stotler and R.E. Stotler.)

Principles of phylogenetic systematics: the species problem. XIII Congreso Mexicano de Botánica. Cuernavaca, Morelos, México. (An invited paper.)

1996. Representing large clades in high-level phylogenetic analyses, with examples from the land plant radiation. Mardi Gras Symposium in Systematics and Evolutionary Biology, Louisiana State University, Baton Rouge. (An invited paper.)

Preliminary results from ITS sequence data analysis in the genus Syntrichia (Musci: Potticeae). Annual meeting, ABLS. Seattle, Washington. (With C. Colacino.)

Influence of mature plants on establishment, and the evolution and ecology of asexual reproduction in mosses. Annual meeting, ABLS. Seattle, Washington. (An invited paper -- with A. E. Newton.)

Biodiversity and the loss of lineages. Conference on: Endangered Languages, Endangered Knowledge, Endangered Environments. Berkeley, California. (An invited paper.)

1997. Molecular systematics of bryophytes. Biennial meeting, International Association of Bryologists. Beijing, China. (An invited paper.)

Systematics Agenda 2000 -- Charting the Biosphere. Annual meeting, Biological Sciences Division, Special Libraries Association. Seattle, Washington, (An invited paper.)

A molecular phylogeny of the bryophytes and their relationships to the tracheophytes based on chloroplast 16s and 23s ribosomal-coding genes. Annual meeting, ABLS. Montreal, Canada. (With J.M.Arrington, L.A. Lewis, R. J. Vilgalys and P.S. Manos.)

Island bryogeography: a case study in the South Pacific. Annual meeting, ABLS. Montreal, Canada. (With D.P. Wall and Mohamed Bin Abdul Majidi.)

Ecological and developmental studies on the dwarf male breeding system of the moss Dicranum scoparium in the North Carolina Piedmont. Annual meeting, ABLS. Montreal, Canada. (With K. Preston.)

Phylogenetic relationships of extant liverworts based on morphological and molecular data. Annual meeting, ABLS. Montreal, Canada. (An invited paper -- with L.A. Lewis, B. Crandall-Stotler and R.E. Stotler.)

Phylogenetic relationships within the Metzgeriidae (simple thalloid liverworts) as inferred from morphological characters. Annual meeting, ABLS. Montreal, Canada. (An invited paper -- with B. Crandall-Stotler and R.E. Stotler.)

An overview of the phylogeny of the mosses inferred from cladistic analysis of morphology and the rbcL gene. Annual meeting, ABLS. Montreal, Canada. (An invited paper -- with D. Gonzalez, J.M. Arrington, L. A. Lewis, and R.J. Vilgalys.)

Phylogeny of the haplolepidous mosses inferred from cladistic analysis of morphology and the rbcL gene. Annual meeting, ABLS. Montreal, Canada. (An invited paper -- with S. Schaffer.)

Major features of the evolution of bryophytes. Annual meeting, ABLS. Montreal, Canada. (An invited paper.)

1998. Compartmentalization in phylogeny reconstruction: philosophy and practice. DIMACS Symposium on Estimating Large Scale Phylogenies. Princeton, New Jersey. (An invited paper -- with P. Soltis and D. Soltis.)

Preliminary analyses of the phylogenetic relationships of Calymperaceae using rbcL sequences. Annual meeting, ABLS. San Juan, Puerto Rico. (With J.A. Wheeler and D.P. Wall.)

Preliminary inferences on the phylogeny of the pleuocarp diplolepidous-alternate mosses from rbcL sequences and morphology. Annual meeting, ABLS. San Juan, Puerto Rico. (With E. De Luna, D. Gonzales, A.E. Newton, and A. Withey.)

Biodiversity: molecular phylogenetics and conservation. 8th Congress of the Federation of Asian and Oceanian Biochemists and Molecular Biologists. Kuala Lumpur, Malaysia. (An invited paper.)

1999. Relationships among museum collections, phylogenetic systematics, and society. Spring Symposium on "Museums, Universities, and Biodiversity in the 21st Century." California Academy of Sciences and Stanford University. (An invited paper.)

An overview of early embryophyte phylogeny: problems and prospects for a "total evidence" solution. Royal Society Discussion Meeting. London, England. (An invited paper.)

Evolution of the major moss lineages. International Botanical Congress. St. Louis, Missouri. (An invited paper -- with A.E. Newton, C.J. Cox, J.G. Duckett, J. Wheeler, B. Goffinet, D. Vitt and T.A.J. Hedderson.)

Phylogenetic relationships within the haplolepideous mosses. International Botanical Congress. St. Louis, Missouri. (An invited paper -- with J. Wheeler, D. Wall, K. Johannes, S. Schaffer, and J. Shaw.)

Phylogeny of desiccation-tolerance in plants. International Botanical Congress. St. Louis, Missouri. (An invited paper -- with Z. Tuba and M.J. Oliver.)

Deep Green: recent results on phylogenetic relationships of the green plants and their evolutionary significance. Keynote Symposium "Phylogeny of Life," International Botanical Congress. St. Louis, Missouri. (An invited paper.)

Early land plant phylogeny, with special reference to bryophytes: problems and prospects for a "total evidence" solution. International Symposium on "Biodiversity and Evolutionary Biology," German Botanical Society. Jena, Germany. (An invited paper.)

Phylogenetics, classification, and species concepts. Annual meeting, Willi Hennig Society. Göttingen, Germany. (An invited paper.)

"Deep Green": recent advances in the phylogenetic relationships of green plants and their evolutionary significance. Annual meeting, Willi Hennig Society. Göttingen, Germany. (An invited paper.)

2000. Deep Green: phylogeny of green plants, comparative genomics, and society. 2000 Genome Symposium, AAAS annual meeting. Washington, DC. (An invited paper.)

Deep Green: lessons learned from collaborative research. Annual meeting, Association of Systematics Collections, Baltimore, Maryland. (An invited paper.)

What is a species? Real world implications of species definitions. Annual meeting, AAAS: Pacific Division. Asland, Oregon. (An invited paper.)

The need for integrated studies of the California flora. Symposium: discovery, communication, and conservation of plant biodiversity in California, Friends of the Jepson Herbarium. Berkeley, California. (An invited paper.)

Spatial distribution of desert bryophyte populations. Annual meeting, ABLS and Botanical Society of America. Portland, Oregon. (A poster -- with M.L. Bonine, L.R. Stark, and D. N. Mcletchie.)

Congruence and convergence in the moss family Calymperaceae: phylogenetic analysis of two chloroplast genes (rbcL and rps4) and morphology. Annual meeting, ABLS and BSA. Portland, Oregon. (With J. A. Wheeler, D. P. Wall, and K. Johannes.)

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