Dean Wm. Taylor
The purpose of this conspectus is to present nomenclatural information about California sedges and to identify UC holdings of type materials. Many North American sedge species were established wholly or in part on material collected by the Geologic Survey of California by William H. Brewer and Henry N. Bolander. The Geological Survey specimens not only served to found the University of California Herbarium, but provided much type material of western plants to Asa Gray and Sereno Watson. Prior to this review, many of the early Carex types in the University Herbarium had not been not identified as type material.
In the era when many sedges were first described, the type concept was not yet fixed. Many species were described from syntypes. Mackenzie, in reviewing protologues of species authored by either Olney or Liberty Hyde Bailey, designated lectotypes using the phrase "is taken as the type." Some workers might consider Mackenzie's choices to be mechanical and thus not in conformance to Article 9A.2, but it should be noted that most recent treatments of Carex (Reznicek and Ball (1980, Standley 1985 etc.) accept Mackenzie's choices, as do I (after all, how can mere botanists counter the choice of a New York lawyer!). I know of no case with respect to California sedges where the mechanical method of selection has resulted in a conflict between the protologue and the historical application of the name.
For ease of reference, numbers used in this conspectus are the Section
or Species numbers as they appear in Mackenzie (1935). The sectional
nomenclature used by Mackenzie is now quite antiquated. Egorova's
monograph [The sedges (Carex L.) 1999) of Russia and adjacent states.
Mo. Bot. Garden Press. 1999] provides needed typification of many supraspecific
taxa used in this list (antiquated names are in RED]
. In a few instances, the protologue was not seen by me (indicated
by Protologue
Not Seen).
Distributional Notes that correct or supplement distribution maps based
on the Jepson Manual are indicated.
A. Subgenus Psyllophora (Degl.) Peterm.
Deutsch. Fl. 602. 1849
( = Primocarex Kuk.,
Das
Pflanzenreich 4, 20 (Heft 38). 1909.)
Spike single, typically androgynous.
About 25 Sections comprising about 75 species of largely extratropical,
typically alpine or arctic, distribution. LECTOTYPE: Carex
pulcharis L.
2. Sect. Capituligerae Kuk. Bot. Jahrb. 27:495. 1899.B. Subg. Carex Terminal spike usually staminate, lateral spikes female, elongate, with a tubular prophyll at base; dorsal suture absent on perigynium. Ca. 60 sections and 1400 species distributed globally. TYPE: Carex hirta L (et generis lectotypus).
( Capitatae Christ. Bull. Soc. Bot. Belgium 24(2):20. 1885.)
SYNOPSIS: 4 species, C. capitata is arctic-alpine of northern Eurasia and North America, extending southward on high mountains to northern Mexico (populations requiring study as subspecifically distinct?); C. rhuiensis Kurtz is from Argentina, the other species, C. oreophila C.A. Mey., is in the mountains of western Asia; C. antarctogena Roivainen of Tierra de Fuego is sometimes separated from C. arctogena. TYPE: Carex capitata L3. Carex capitata L. ssp. arctogena (H. Smith) Boecher3. Sect. Inflatae Kuk.
Arctic. Alp. Res. 3, 2:144. 1971. |Jepson Manual Distribution |Specimens|
TYPE: based on C. arctogena H. Smith
NOTE: a special status plant in Oregon.Carex arctogena H. Smith
Acta Phytogeogr. Suec. 13:193. 1940.
TYPE: Swedish Lapland, H. Smith s.n. (S, DAO)
Das Pflanzenreich 4, 20 (Heft 38):96. 1909.
SYNOPSIS: two to perhaps four dry site alpine taxa of western North America, o in the Sierra Nevada5. Carex breweri F. Boott var. breweri25. Sect. Callistachys (Heuffel) Mack.
Illus. Carex. p. 142 & pl. 455. 1867. | Jepson Manual Distribution |Specimens|
TYPE: California, Siskiyou Co., Mt. Shasta W.H. Brewer 1422 in 1863 (isotype GH27152). No type material at UC. The relationships of Carex breweri var. paddoensis (Suksd.) Cronq. (a rare plant in Idaho, isotype NY11263) and Carex engelmanii L.H. Bailey warrant study.5a. Carex subnigricans Stacey
Leaf. Western Bot. 2:167. 1939. |Jepson Manual Distribution |Specimens|
TYPE: California, Tuolumne Co., Mt. Dana, J.T. Howell 14519 (holotype CAS259816, isotypes, GH27450, US1765699, UC618535). Paratypes cited in the protologue include F.W. Peirson 10828, 11400, 12280 (CAS, RSA), McCraken s.n. July 25, 1936 (CAS), C. Sharsmith 102 (CAS), 2356 (CAS, isoparatype UC714264), D.D. Keck 4657, 4914 (CAS, DAO) J.T. Howell 14136, 14154 (CAS); H.S. Yates 6012 (VTM, not at UC). The UC duplicates of the paratypes were not cited in the protologue, and are therefore isoparatypes (paratypes are not listed in the online CAS type catalog).
NOTE: a special status plant in Oregon.Carex rachillis B. Maguire
Brittonia 5:199. 1941
TYPE: Utah, Summit County, Gilbert Peak, B. Maguire et al. 14668
N. Amer. Flora 18, 1:26. 1931.
SYNOPSIS: 5 dry ground, calcareous, arctic-alpine or alpine, coastal species in the western part of North America north of Mexico, with C. nigricans in North America, C. micropodioides V. Krecz in southern Europe, C. micropoda of boreal Europe and Asia, and Alaska, and Carex cephalotes F. Muell. is found in New Zealand.9. Carex nigricans C.A. Mey. |Jepson Manual Distribution |Specimens|22. Sect. Polytrichoideae Tuck.
Mem. Acad. Imp. Sci. St.-Petersburg. Ser. 6, Sci. Math. 1:211 & pl. 7. 1831.
TYPE: from Unalaska, Alaska, Ludolf Adelbert von Chamisso (isotype GH27346). Two specimens in the NY catalog that are listed as types (NY11234 & NY11224) are likely not such. Egorova (1999) lists the type as Unalaschka, Mertens (Holotype LE, isotypes LE), casting into doubt the citation of a Chamisso specimen as the type.
Enum. Caric. 8. 1843.
SYNOPSIS: Monotypic, Sphagnum bogs in North America (very rare in California, known from NCo bogs).198. Carex leptalea Wahl. - | Jepson Manual Distribution |Specimens|24. Sect. Filifoliae Tuck.
Kongl. Vetensk, Acad. Nya Handl. 24:139. 1803.
TYPE: "in America borealis, secundum herbarium Cl. Torneri".
NOTE: A special status plant in California (CNPS List 2, vouchered from Del Norte, Humboldt, Marin [extirpated] and Trinity counties), Idaho, and Oregon.
Enum. Caric. 8. 1843.
SYNOPSIS: 6 dry ground, montane species of western North America, with C. arsenii Kuk. in the Michoacan subalpine;
C. longissima M.E.Jones, of the Sierra de la Laguna, Baja California isolated and reclictual.205. Carex filifolia Nutt.- | Jepson Manual Distribution |Specimens|28. Sect. Scripinae Tuck.
Gen. N. Amer. Pl. 2:204. 1818.
TYPE: Th. Nuttall, on the dry plains and gravely hills of the Missouri, in 1813.
DISTRIBUTIONAL NOTE: the UC-JEPS specimen records are not yet updated. An additional CA station is: Shasta Co., Burney Spring, DWT 9839.207. Carex exserta Mack.
Bull. Torrey Bot. Club. 42: 620. 1915. | Jepson Manual Distribution |Specimens|
TYPE: based on C. filifolia var. erostrata Kuk.Carex filifolia var. erostrata Kuk. in Engler
Das Pflanzenreich 4, 20 (Heft 38):86. 1909.
LECTOTYPE: Ezra Brainerd 111. Lectotype designated by Mackenzie (Erythea 8:52. 1922); in North Am. Flora 18:180. 1931, this specimen is cited as Brainerd, July 11, 1897 (lectotype GH27229, isolectotype UC864).
SYNTYPES: Hall et Chandler 616 (UC865), M.E. Jones 2908 (UC825102)
Enum. Caric. 8. 1843.
SYNOPSIS: 7 species in a polymorphic, circumboreal complex, often of calcareous dry soils: in North America, one into Eurasia. Starr et al. (AJB 86:563-577. 1999) found this Section was misplaced within Subg. Psyllophora [Primocarex]. Note that Mackenzie, although he did not treat subgenera, placed the Scripinae next to Sect. Digitatae, which is Subg. Carex. Following Mackenzie's placement has the utility of making Subg. Psyllophora [Primocarex] monophylletic.240. Carex pseudoscirpoidea Rydb.31. Sect. Triquetrae Carey ex Olney
Mem. N.Y. Bot. Gard. 1:78. 1900. | Jepson Manual Distribution |Specimens|
LECTOTYPE: Wyoming, Spanish Peak, Yellowstone Park, Rydberg 3064 (NY11290) Lectotype designated by Mackenzie N.Amer. Flora 18:209. 1935, and accepted by Dunlop (Novon 7:355-356, 1997).
SYNTYPE: Rydberg 3412 (NY214536)
NOTE: a special status plant in California (CNPS List 2, vouchered from Alpine, Inyo and Mono counties)Carex scirpoidea Michx. var. pseudoscirpoidea (Rydb.) Cronq.242. Carex scabriscula Mack.
Univ. Wash. Publ. Biol. Sci. 17(1):325. 1969.Carex scirpoidea Michx. ssp. pseudoscirpoidea (Rydb.) D.A. Dunlop
Novon 7:355. 1997.
Bull. Torrey Bot. Club 35:268. 1908.
TYPE: Oregon, Cascade Mountains, Cusick 2849 June 30, 1902 (holotype NY11317, distributed as C. feta, isotypes DS490735, CU, ORE17812, OSC153663, POM, US528631, WS).241. Carex gigas (Holm) Mack.
Bull. Torrey Bot. Club. 35:268. 1908. | Jepson Manual Distribution |Specimens
TYPE: based on C. scirpoidea var. gigas Holm
NOTE: a special status plant in Oregon.Carex scirpoidea var. gigas Holm
Am. Jour. Sci. Series 4, 18:20 & f. 8. 1904.
TYPE: Dunlop (1997) states that Holm failed to designate a type, and that Mackenzie's reference to a type from Siskiyou County, California, C.G. Pringle s.n., August 18, 1881 (NY11321) did not constitute a lectotypification. She, however, did not select an alternate lectotype. The disjunct distribution and ecological differences may indeed warrant recognition of C. gigas as a valid taxon.
Proc. Am. Acad. 7:395. 1868.
SYNOPSIS: 5 dry ground species in temperate North America.252. Carex triquetra F. Boott | Jepson Manual Distribution |Specimens|33. Sect. Firmiculmes Kuk.
Trans. Linn. Soc. 20:126. 1846. Protologue Not Seen
TYPE: habitat in California, T. Nuttall 23 (K?, possible type NY11386).Carex monticola Dewey in Torr.
Botany Mexican Boundary Survey, 229. 1859.
TYPE: California, San Diego County, mountains east of San Diego, C.C. Parry s.n. in 1850 (possible type NY11230)
Das Pflanzenreich 4, 20 (Heft 38):93. 1909.
SYNOPSIS: 3 dryland species in western North America, all found within California, two endemic to the CA-FP; the reported isolated station for C. multicaulis in Elko County, Nevada (Intermountain Flora, Cronquist et al. 1977) requires taxonomic study.257. Carex multicaulis L.H. Bailey
Bot. Gaz. 9:118. 1884. | Jepson Manual Distribution |Specimens|
LECTOTYPE: California, Mariposa County, Yosemite Valley, J. Torrey 544 (NY). The lectotype was designated by Mackenzie (Erythea 8:51. 1922); there are NY11231, NY11232, NY11233 with this collection number, as well as GH58547 .
SYNTYPES: California, Mendocino County, Ukiah, H.N. Bolander 39; Mariposa Grove, W.H. Brewer 1635 [UC8000] & [UC924], Dr. W. Hillebrand 2306; Plumas County, Mrs. Ames s.n.; Duffield's Ranch, J.M. Bigelow s.n.; Oregon, Almanden, Thos. Howell s.n. in 1884
NOTE: UC1119 is another sheet that might be type material, but it is labeled only as "State Survey" without collector or collection number being noted on the sheet.257a. Carex tompkinsii J. Howell
Leaf. Western Bot. 9:185. 1961. | Jepson Manual Distribution |Specimens|
TYPE: California, Fresno County, Copper Creek trail, Kings Canyon, J.T. Howell 35333, June 6, 1960 (holotype CAS428953, isotype CAS429306, isotypes US2604279 & US2853449, NY11379, WTU262400, DAO OSC132834, TEX). Note that the WTU specimen is listed as the holotype, contrary to the protologue, and is an isotype.
NOTE: A special status plant in California (CNPS List 4), vouchered from Mariposa and Fresno counties.259. Carex geyeri F. Boott | Jepson Manual Distribution |Specimens|
Trans. Linn. Soc. 20:118. 1846. Protologue Not Seen
TYPE: Rocky Mountains, C.A. Geyer 332 (holotype K?, probable isotypes GH58546, NY11120)
NOTE: Special status plant in British Columbia and California (CNPS List 4, vouchered from Butte, Humboldt, Plumas, Siskiyou and Trinity counties).
DISTRIBUTIONAL NOTE: the stated Jepson Manual range does not reflect the nSN stations.
56. Sect. Anomalae Carey in A. Gray
Gray's Manual 557. 1848.
SYNOPSIS: a section of about 30 species strongly developed in eastern Asia; several species in Australasia, with 2 in North America, C. amplifolia in the west and C. scabrata in the eastern U.S. LECTOTYPE: C. scabrata Schwein398. Carex amplifolia F. Boott in Hooker60. Sect. Microrhynchae Drej. ex L.H. Bailey (Proc. Amer. Acad. Arts. Sci. 22:76)
Fl. Boreali-Amer. 2:228 & pl. 226. 1839. | Jepson Manual Distribution |Specimens|Perigynia|
TYPE: Columbia River, David Douglas (holotype presumably at K)
DISTRIBUTIONAL NOTE: the stated Jepson Manual range omits reference to the Montana stations, where this taxon is considered a special status plant, and New Mexico (Roalson, E.H., Jones, S.D., Allred, K.W. Sida 16(3): 592-594. 1995). Special status plant in British Columbia.
(= Sect. Atratae Kunth. Enum. Pl. 2:431. 1837.)
SYNOPSIS: a section containing many species and strongly developed in the Holarctic, particularly in North America and Asia. One species occurs in the Andes (C. atropicta Steid); one is reported from Australia (Mackenzie considered erroneous). Mackenzie treated 29 species in North America. TYPE: Carex atrata L. (Europe) Egorova treats 8 Subsections in Russia.411. Carex parryana Dewey ssp. hallii (Olney) Murray
Brittonia 21:71. 1969. | Jepson Manual Distribution |Specimens|
TYPE: based on Carex hallii Olney
NOTE: UC1736037 not yet in the herbarium database, see Madrono 35: 164-166. (1988). A special status plant in CA (CNPS List 2).Carex parryana var. hallii (Olney) Kuk. in Engler
Das Pflanzenreich 4 (Heft 38), 20:388. 1909.
TYPE: based on C. hallii OlneyCarex hallii Olney in Hayden413. Carex norvegica Retz. ssp. norvegica (Subsection Alpinae Kalela ex Egor.)
Rep. U.S. Geol. Survey 5:496. 1872.
LECTOTYPE: Hayden s.n, June 1871, Rocky Mountains (NY, but not in types catalog).
Lectotype designated by Murray 1969. Brittonia 21:71.(and previously by Mackenzie 1935)
SYNTYPES: E.M. Hall & J.P.Harbour 617 (US29651, GH27253, LCU, NY11134), T.C.Porter (NY11135).
Florae Scandinaviae Prodromus 179. 1779. | Jepson Manual Distribution |Specimens|
TYPE: Norway. ? LD
NOTE: UC1736045, UC1736056 are not yet in SMASH, see Madrono, 37: 64-65. (1990).
A special status plant in California (vouchered only from the White Mountains, Mono County) and Oregon.417. Carex spectabilis Dewey | Jepson Manual Distribution |Specimens|
Am. J. Sci. 29:248, pl. X, f. 76. 1836.
TYPE: "found in Arctic regions" NY11336 is a possible typeCarex invisa L.H. Bailey420. Carex raynoldsii Dewey | Jepson Manual Distribution |Specimens|
Proc. Amer. Acad. 22:82. 1886.
TYPE: "Summit Camp, California, Kellogg, the type growing ‘in exceedingly tough and
matted clumps". Holotype presumably at BH.
TYPIFICATION: Bailey’s exact wording of the protologue suggests by implication that he took the Kellogg collection as the holotype, and the other specimens cited were representative material (paratypes). Throughout Bailey’s Carex synopsis, he did not make specific reference to herbaria in which type material was seen. He did note "if I have seen and examined critical or historic specimens, the collector’s name is printed in Italics", as is the case in this instance. PARATYPES: Ebbett’s Pass, W.H.Brewer 2084 (UC993) and 2076 (UC994); Big Trees, Hillebrand; Lassen’s Peak above snow, W.H.Brewer 2186 (UC992) ["a globular spiked form"]; Carson Pass, W.H. Brewer 2126 (UC999); Selkirk Range, British Columbia, Macoun." The following specimen listed as a syntype: California, Mariposa County, Yosemite Valley, H.N. Bolander 6198 (US30161), but this number at other herbaria also serves as type material of C. whitneyi and C. paucicostata, and was not mentioned in the protologue.Carex jepsonii Mackenzie ined (not Carex jepsonii mackenzii as in the NY database?)
an ined. name attributed to NY11184 as a type in their database, W.L. Jepson 4577, 31 July 1911. The specimen label is in Jepson's handwriting, and the specimen appears (from my inspection of the image) to be Carex spectabilis Dewey.a name possibly referable to this taxon:
Carex tolmiei var. invisa Kuk. 1909 p. 412
Am. J. Sci. Series 2, 32:39. 1861.
LECTOTYPE: Idaho, Fremont Co., Pierre's Hole, Snake River, June 20, 1860, V.F. Haden GH27389
Lectotype designated by Murray (Brittonia 21:71. 1969). The GH specimen catalog lists the specimen as an isolectotype426. Carex helleri Mack. | Jepson Manual Distribution |Specimens|
Erythea 8:80. 1922.
TYPE: Nevada, Washoe County, Mt. Rose, A.A. Heller 9975 (isotype US509004, NY11145).427. Carex epapillosa Mack. in Rydb.
Fl. Rocky Mtns. 138, 1060. 1917. | Jepson Manual Distribution |Specimens|
TYPE: Utah, Marysville, M.E. Jones 5345 (holotype NY11098, isotype NY11099, RM, US270933). According to Lenz (1986), Jones was there on July 30th et seq, 1894, and nos. 5846-47c on August 21st "at Marysville"
NOTE: Special status plant in British Columbia.Carex heteroneura W. Boott var. epapillosa (Mackenzie) F.J. Herm.428. Carex heteroneura W. Boott in S. Watson
Rhodora 71:428. 1968.
Bot. Calif. 2:240. 1880. | Jepson Manual Distribution |Specimens|
TYPE: "In the Sierra Nevada; Lake Tahoe to Bear Valley, A. Kellogg s.n., August 3 (holotype GH27259, possible isotype US28206).Carex atrata var. erecta W. Boott in S. Watson
Bot. Calif. 2:239. 1880.
TYPE: unknown.
PROTOLOGUE: "in the Sierra Nevada, not rare, from 5000 ft. (Big Trees, Bolander) to 10,000 or 12,000 ft altitude both the typical form and the varieties". At UC, there are apparently no sheets that correspond to the appropriate Bolander collection.Carex quadrifida L.H. Bailey
Proc. Calif. Acad. Series II, 3:104. 1891.
SYNTYPES: California, Mt. Dana, H.N. Bolander 5046 (UC623 & UC72441,
US319251, US2012647, US469687, NY11295), W.H. Brewer 1773 (UC620, UC8076); DS55002 is listed as isotype in their catalog, but this is rather a syntype.Carex quadrifida var. lenis L.H. Bailey
Proc. Calif. Acad. Series II, 3:105. 1891.
SYNTYPES: Bolander 5046 (UC623 & UC72441, NY11296), Kellogg & Harford 1080 [US28217, NY232332], Kellogg from Donner [which in fact may be the former specimens].Carex quadrifolia var. caeca L.H. Bailey
Bot. Gaz. 21:8. 1896.
TYPE: "San Diego County", Tahquitz Meadow, San Jacinto Mountain, 8000 ft altitude, H.E. Hasse July 1892 (holotype BH?).
NOTE: the actual type location is in Riverside County [ca. 33.76820 -116.66362]Carex atrata var. discolor L.H. Bailey
London Jour. Bot. 26:321. 1888.
non: Smiley Univ. Calif. Pubs. Bot. 9:124. 1921.
TYPE: no specimen designated
PROTOLOGUE: "mountains of Colorado and Utah and southward"430. Carex albonigra Mack. in Rydb.
Fl. Rocky Mts. 137. 1917. | Jepson Manual Distribution |Specimens|
TYPE: Wyoming, Needle Mountain, Merritt Cary 613 (US858947).
PROTOLOGUE:"Mountain meadows: Mont.-Colo.-Ariz.-Utah. Subalp.-Alp."
Murray (1969) did not review the UC material.435. Carex mertensii Prescott - | Jepson Manual Distribution |Specimens|
Bong. Mem. Acad. St.-Petersburg VI. 2:168. 1832.
TYPE: Alaska, Sitka, MertensCarex columbiana Dewey
Am. J. Sci. 30:62. 1836.
TYPE: Columbia River, Scouler (NY11039)436. Carex serratodens W. Boott in S. Watson
Bot. Calif. 2:245. 1880. | Jepson Manual Distribution |Specimens|
TYPE: no specimens cited at publication.
PROTOLOGUE: California, 'in herb Olney, but locality or collector not stated' The text is Watson's attempt to account for the absence of a citation of any of the original material. A type fragment, without collector or locality, is at GH57539.61. Sect. Phacocystis Durmot. Fl. Belg. 146.Carex bifida Boott ex Olney
Proc. Amer. Acad. 7:394. 1868.
non C. bifida Roth in Steud. Nom. ed. 1 155, 1821.
SYNTYPES: California, Monterey County, Salinas Valley, W.H. Brewer 574 (UC661); and, Mendocino County, Red Mountain, H.N. Bolander 6476 (UC8043, GH57535).Carex aequa C.B. Clarke
Kew Bull. Add. Ser. 8:86. 1908.
TYPE: California, San Mateo County, Crystal Springs Lake, E.L. Greene s.n., distributed as C.F. Baker 811 (holotype K, isotypes GH27126, UC143733, NY38978)
(Sect. Acutae (Fries) H. Christ, Fl. Scan. 191. 1835)
SYNOPSIS: a group of ca. 80 species of swamps of the Holoarctic; probably best developed in Asia. Mackenzie recognized 42 species in North America. Several species in southern South America and Australasia. Barely reaching North Africa. Mackenzie notes "the fundamental distinguishing characters are found in the underground parts; these are unfortunately usually not gathered by collectors (Egorova 1999 also laments incomplete specimens). The scales and spikes vary much in size and shape in the same species, and distinctions based upon them need an unusual amount of verification by a study of abundant material". Partially fertile hybrids complicate identification. TYPE: Carex cespitosa L. (Europe). Egorova (1999) places this Section in Subg. Kreczetoviczia Egor, along with Sect. Temnemis (below)and 6 other sections [Abditispicae Wheeler, Graciles (Tuck. ex Kuk.) Ohwi; Forficulae Franch. ex Raymond; Praelongae (Kuk.) Nelmes and Tuminescens Y.I. Chang & Y.L. Yang)]Subsection Rigidae Fries ex Kuk. Pflanzenr. 38:229. 1909 TYPE: Carex rigida of Eurasia.
442. Carex scopulorum Holm var. bracteosa (L.H.Bailey) F.J. Herm.
Leaf. W. Bot. 9:16. 1959.| Jepson Manual Distribution |Specimens|
TYPE: based on Carex vulgaris var. bracteosa BaileyCarex vulgaris var. bracteosa L.H. Bailey
Proc. Amer. Acad. 22:81. 1887
TYPE: California, Alpine County, by a lake at Ebbett’s Pass, W.H. Brewer 2015, August 3, 1863 (holotype BH?, isotypes DS49838, US469730, US319238 & US319237, GH27481, GH27482, UC1509 & UC8026)Carex rigida var. hesparia C.V. Piper
Cont. U.S. Natl. Herb. 11:173. 1906.
TYPE: " Ebbett’s pass, California, alt. 8,000 feet Collected by Brewer."Subsection Vulgares Aschers.Carex gymnoclada Holm
Amer. J. Sci. Series 4, 14:424. 1902.
TYPE: Oregon, Wallowa County, bogs of Hurricane Creek, W. Cusick 2487 (holotype LCU?, isotype US385394, ORE16961, GH27249, GH27250, NY11131)
Fl. Brandenb. 1:774. 1864. TYPE: Carex vulgaris Fries (Europe)452. Carex nebrascensis Dewey | Jepson Manual Distribution |Specimens|
Am. J. Sci. Series 2:18. 102. 1854.
TYPE: Nebraska, F. V. Hayden s.n. in 1853 (holotype GH27339, isotypes NY11235, NY11234, possibly NY11262, MO)Subsection Aquiticae Egor. Bot. Zurnal 75, 6: 864. 1990. TYPE: Carex aquatilis Wahl.Carex jacintoensis Parish
Bull. S. Cal. Acad. 4:110 & pl. 16. 1905.
SYNTYPES: "Collected at high altitudes in the San Jacinto Mts., July and Aug, 1901, by Mr. H.M. Hall, 2461, Tahquitz Valley, 7000 ft. alt., 2561 [UC72389], Fuller’s Mill, 8,800 ft. alt. 2483, 2484, Tamarack Valley, 9,000 ft alt.; 2338 [UC961], Deep Spring, 9000 ft. alt. In Mr. Hall’s Botanical Survey of the San Jacinto Mt. These specimens are referred to C. Hallii, C. Nebraskensis and C. nudata." This synonym was not attributed to C. nebrascensis by Standley (Sys. Bot. Monogr. 7:73. 1985). DS78003 is labeled as the holotype.456. Carex aquatilis Wahl. var. dives (Holm) Kuk.
Das Pflanzenreich 4, 20 (Heft 38):311. 1909.| Jepson Manual Distribution |Specimens|
TYPE: based on C. dives Holm.Carex dives Holm.
Amer. J. Sci. Arts., ser. iv, 17:312. 1904
LECTOTYPE: Oregon, Henderson s.n. (US85957, isolectotype OSC1929). Standley (Sys. Bot. Monogr. 7:67. 1985. The authority citation for this combination, (Holm) L. Standley in the Jepson Manual is in error.
SYNTYPES: "Collected in Oregon by Mr. L.F.Henderson; in California" ‘in open swamps with C. utriculata, 12 mile house San Jose R.R.’ collected by H.N.Bolander, and in Chilliwack Valley, British Columbia by Mr. James M. Macoun (NY11076)." The latter syntype is listed as a lectotype in the NY catalog and annotated as such by L. Standley in 1981, but not published as such.457. Carex aquatilis Wahl. var. aquatilis| Jepson Manual Distribution |Specimens|
Kongl. Vetensk. Acad. Nya Handl. 24:165. 1803.
TYPE" Lapland, Wahlenberg s.n. (C).449. Carex lenticularis Michx. var. lipocarpa (Holm) L. Standley
Sys. Bot. Monogr. 7:67. 1985. | Jepson Manual Distribution | Specimens |
TYPE: based on Carex vulgaris var. lipocarpa HolmCarex vulgaris var. lipocarpa Holm
Amer. J. Sci. Series 4, 17:308. 1904.
PROTOLOGUE: "Collected in Alaska at several stations [cf. US85936], on Vancouver Island, in the Selkirk Mountains, British Columbia, and in the Chilliwack Valley [US3151517], by Mr. James M. Macoun and others."
LECTOTYPE: British Columbia, J.M. Macoun 33360 (lectotype LCU, isolectotypes GH27484 NY11076, US3151517). On the NY11076 sheet, the annotation label lists the specimen as an isolectotype of C. dives, which is not consistent with the publication - perhaps a misplaced annotation label (sheet NY11076 is no longer in their database as of December, 2000).Carex kelloggii W. Boott in S. Watson
Bot. Calif. 2:240. 1880.
TYPE: California, Sierra Nevada, Lake Tahoe to Bear Valley, A. Kellogg s.n. (GH27287). Lectotype designated by L. Standley, Sys. Bot. Monogr. 7:67. 1985. NOTE: as of 3/19/99 the GH specimen is listed as an isosyntype in their catalog.
SYNTYPES: "Lemmon; and Alta, Wasatch Mountains, Utah, M.E. Jones", as given in the protologue. Lenz (1986) states that M.E.Jones was in Alta in 1879, on July 29th-August 13th.448. Carex lenticularis Micha. var. impressa (L.H.Bailey) L. Standley
Sys. Bot. Monogr. 7:65. 1985.| Jepson Manual Distribution | Specimens |
TYPE: based on C. interrupta var. impressa L.H. BaileyCarex interrupta var. impressa L.H. Bailey
Mem. Torrey Bot. Club 1:18. 1889.
LECTOTYPE: California, Summit Camp, A. Kellogg s.n. (lectotype BH, isolectotype GH27283). Lectotype by L. Standley Sys. Bot. Monogr. 7:65. 1985. NOTE: as of 3/19/99 the GH specimen is listed as an isosyntype in their catalog.
SYNTYPE: Yosemite Valley, H.N. Bolander 6198 (UC981, UC1019 US30161) but Bolander specimens with this same number are also type material of C. whitneyi, C. paucicostata)448. Carex paucicostataMack.
Erythea 8:74. 1922.
LECTOTYPE: "Summit Camp, California (Kellogg)" (lectotype BH, isolectotype GH27283). Lectotype by L. Standley Sys. Bot. Monogr. 7:65. 1985.
PARATYPES: Yosemite Valley, H.N. Bolander 6198; Lake Tenaya, J.W. Congdon (UC115763); Crescent Lake, Mariposa Co., J.W. Congdon; Hocket Meadow, Tulare Co., W.R.Dudley 1006 (in part); Kern Lake, Tulare Co., W.R.Dudley 2044; Little Lake, El Dorado Co., E. Brainerd 76 (MID?); Strawberry Creek, El Dorado Co., E. Brainerd 35 (MID?); Truckee, Hitchcock 258; Stanislaus Forest, Alpine Co., Eggleston 9517; shore of Silver Lake, Amador Co., W.H. Brewer 2099 [UC1050]; South Fork San Joaquin River, H.M. Hall & H.P. Chandler 651 (UC1143); Harmon Meadow, Tulare Co., W.R.Dudley 2819.450. Carex lenticularis var. limophila (Holm) Cronq.
|Jepson Manual Distribution|Specimens|
Vascular Plants of the Pacific Northwest 1:279. 1069.
TYPE: based on C. vulgaris var. limophila HolmCarex vulgaris var. limophila Holm.
Amer. J. Sci. Arts Ser. IV, 17:307. 1904.
LECTOTYPE: Alaska, Bering Sea, St. Paul Island, J.M.Macoun 16613 (Lectotype K, isolectotype GH27484). Lectotype designated by Standley op. cit. p. 66.Subsection SentaeCarex hindsii C.B. Clarke
Kew Bull. Additional Series 9:70. 1908.
LECTOTYPE: Columbia River, Hinds s.n. (Lectotype K, isolectotype GH27260). Lectotype designated by Standley, op. cit. p. 66.
SYNTYPE: "Montana, alt. 2250 m, Rydberg et Bessey, n. 3740" (presumably at K).459. Carex barbarae Dewey in Torr. | Jepson Manual Distribution | Specimens |
Bot. Mexican Boundary Survey 231. 1859.
TYPE: "banks of streams, Santa Barbara, California; C.C.Parry." (holotype GH27522).
NOTE: a special status plant in Oregon.Carex wilksesii Torr.
U.S. Explor. Exp. 17:477. pl. 17. 1874. Protologue Not Seen
TYPE: banks of the Sacramento" No specimen listed in NY catalog.Carex laciniata W. Boott
Ill. Gen. Carex 4:175. Pl. 594. 1867.
TYPE: "American Fork of the Sacramento, W. Rich" (NY11187)Carex lacunarum Holm.
Amer. J. Sci. Series 4, 17:316. f 12-13, 1904
SYNTYPES: "California, lagoon at Sebastopol, Sonoma County, collected by Mr. A.A. Heller (No. 5797) (syntypes US430229, NY11188) [July 1, 1902], and in a marsh at Berkeley by Mr. J. Burtt Davy". No Davy material at UC was found that might be original material.Carex senta Boott x Carex barbarae
Hansen 641, Jackson, Amador County, (Herb. Columbia Coll, not in NY types catalog.)461. Carex senta Boott | Jepson Manual Distribution | Specimens |
Illus. Carex 174. 1867.
TYPE: California, Santa Barbara Co., Santa Ynez Mountains, W.H. Brewer 350 in 1864 (2 type sheets, GH27430 & GH27431, no indication as to which is holotype)Carex auriculata L.H. Bailey
Mem. Torrey Bot. Club. 1:19. 1889.
non C. auriculata Franch., 1886.
TYPE: "California, Colma, N.J. Anderson 1852. Hb. Holm."
PARATYPES: Clark's Ranch, Yosemite Valley, Henry N. Bolander 6208 and 6212. Bolander 6212 (UC877) is a specimen of C. lanuginosa Michx.Carex austromontana Parish
Bull. S. Cal. Acad. 4:108 & pl. 15. 1905.
TYPE: holotype DS489410
PARATYPES: "growing in tussocks on stream banks in the higher mountains. San Bernardino Mts. Mill Creek 6,000 ft. alt., 2485 Parish, July, 1892, type; 5031 Parish, June 1901. Bear Valley, 6,500 ft. alt., 3279 Parish; June, 1984. San Jacinto Mts., Round Valley, 9,500 ft alt., Dr. Hasse, June 3, 1903."Carex bishalii C.B. Clarke
Kew Bull. Add. Ser. 8:70. 1908.
TYPE: California, Mariposa County, Yosemite, W.H. Brewer 1648 (holotype K according. Standley).
NOTE: listed as a synonym of C. senta by Mackenzie (Fl. North America 18:400. 1935), and as synonym of C. nudata by Standley). NY25130, J.Torrey s.n. in 1865 from Clark's Ranch (now Wawona) is listed as possible type, but this specimen is not mentioned in the protologue.Carex bolanderi Gand.
Bull. Bot. Soc. France 66:296. 1920. Protologue Not Seen QK1 S4 or A.455
TYPE: from California.
not C. bolanderi Olney, 1868.460. Carex schottii Dewey in Torr. | Jepson Manual Distribution | Specimens |Subsection Strictae
Bot. Mexican Boundary Survey 231. 1859.
TYPE: "Banks of rivers, Santa Barbara, California; C.C.Parry"468. Carex nudata W. Boott in S. Watson | Jepson Manual Distribution | Specimens |
Bot. Calif. 2:241. 1880.
TYPE: California, Marin County, H.N. Bolander 2299 (lectotype GH27349, isolectotypes NY WS ). Lectotype designated by Standley?? (Sys. Bot. Monogr. 7:76.) NOTE: as of 3/20/00 the GH specimen is listed as an isosyntype in their catalog.
SYNTYPES: Bolander 3836, 4638, 6202. Bolander 121 listed by Mackenzie (Fl. N. Amer. 18:407. 1935) was not cited in the protologue, but this number may be the corresponding number in Bolander's' Catalog of Flora of San Francisco".Subsection Forsiculae Franch ex Kuk. Pflanzenr. 38:334. 1909Carex nudata f. sessiliflora Kuk.
Das Pflanzenreich 4, 20 (Heft 38):337. 1909.
LECTOTYPE: California, Amador County, G. Hansen 636. Lectotype designated by Standley (Sys. Bot. Monogr. 7:76.). Mackenzie (N. Amer. Fl. 18:401. 1935) made this a synonym of C. senta Boott.
TYPE: Carex forficula Franch & Savant; Egorova (1999) however does not admit C. angustata to this subsection.471. Carex angustata F. Boott| Jepson Manual Distribution | Specimens |
Fl. Bor.-Amer. 2:218. 1839.
LECTOTYPE: Columbia River, Scouler (K), designated by Standley (Sys. Bot. Monogr. 7:47.)
NOTE: formerly a special status plant in Idaho.62. Sect. Temnemis (Rafin.) V. Krecz.Carex eurycarpa Holm.
Am. J. Sci. Arts. Series 4, 20:303. 1905.
TYPE: Washington, Klickitat County, meadows near the Columbia, June 1885, W. Suksdorf 816 (holotype LCU, isotypes GH27221, US85957).
(= Sect. Cryptocarpae Tuck. Enum. Caric. 11. 1843
SYNOPSIS: perhaps only a subsection of Acutae, includes 15 saline or marsh species in the Pacific and Atlantic. Standley showed several characters of leaf anatomy shared by C. obnupta and C. lyngbyei not shared by Acutae, but C. barbarae is intermediate between the two sections. A plant from serpentine in the southern Sierra Nevada foothills perhaps represents an undescribed species.481. Carex obnupta L.H.Bailey | Jepson Manual Distribution | Specimens |
Proc. Calif. Acad. Sci. Series 2, 3:104. 1891.
LECTOTYPE: California, San Mateo County, San Mateo, A. Kellogg s.n. (BH, isolectotype DS54912). Mackenzie (Erythea 8:81. 1922) designated this specimen as the ‘type’. In this instance, Mackenzie’s choice was not exactly mechanical, being that he noted that the Kellogg & Brannan specimen from Donner Pass was another taxon.
SYNTYPES: Sierra Nevada (Donner), Kellogg & Brannan; Fort Point, San Francisco, H.N. Bolander 2302 (UC1309, labeled as near San Francisco Bay, is a possible syntype).477. Carex lyngbyei Hornem. ssp. cryptocarpa (C.A. Mey) Hult.27. Sect. Acrocystis Dumort.
Jepson Manual Distribution | Specimens |
Flora Kamtch. 1:188. 1927.
TYPE: based on Carex cryptocarpa C.A.Mey.
NOTE: the Jepson Manual does not treat this subspecies; Carex lyngbyei ssp. lyngbyei is confined to northern Atlantic Ocean shores. A special status plant in California (CNPS List 2, vouchered from Humboldt, Mendocino and Marin counties only).Carex cryptocarpa C.A. Mey.
Mem. Sav. Etr. Petersb. 1:226. 1831
TYPE: "Habitat in Unalaschka et Kamtschatka", Eschscholz (Holotype LE, isotypes LE(3)
Sect. Montanae (Freis.) H. Christ Fl. Scan. 188. 1835.
SYNOPSIS: a group of many dry ground species in Eurasia and North America, with about 30 species; one known to Mackenzie in the Andes, and one in the Azores. Not represented in Australasia. In California, we perhaps have one described taxon from gabbro in the northern Sierra Nevada foothills (Pine Hill), and perhaps a second in nSNH (passing as C. rossii). North American members are under study.222. Carex inops L.H.Bailey | Jepson Manual Distribution | Specimens |
Proc. Am. Acad. 22:126. 1886.
TYPE: Oregon, sandy grounds on subalpine slopes of Mt. Hood, L.F. Henderson s.n., July 1884 (BH?, isotypes GH27273 CAS203910, ORE17107, NY11164).
NOTE: a special status plant in California (CNPS List 2, vouchered from Plumas and Siskiyou counties only). The printed Jepson Manual distributional statement omits a nSNH record.Carex pensylvanica var. vespertina L.H.Bailey
Mem. Torrey Bot. Club 1:74. 1889
SYNTYPES: Cascades of the Columbia, Oregon, T. Howell (ORE?); east Portland, L.F.Henderson 1782 (GH27369, ORE?, US29709); Mt. Adams, T. Howell (ORE?); Vancouver Island, J.M. Macoun (UC?)225. Carex globosa W. Boott| Jepson Manual Distribution | Specimens |
Proc. Linn. Soc. 1:259. 1845
Trans. Linn Soc. 20:125. 1846.
TYPE: California, Nuttall (holotype K?)
DISTRIBUTIONAL NOTE: the Jepson Manual distributional statement does not reflect a single n Baja CA
station (Cerro Blanco, Moran 17592, as cited in Hermann, USDA Agricultural Handbook 467 1974)Carex umbellata var. globosa Kuk. in Engler
Das Pflanzenreich 4, 20 (Heft 38):453. 1909.226. Carex brainerdii Mack.| Jepson Manual Distribution | Specimens |
Bull. Torrey Bot. Club. 40:534. 1913.
TYPE: California, El Dorado County, Sierra Nevada range, mountain north of Slippery Ford., Ezra Brainard 121, 19 July 1897 (holotype, ‘herb. Brainerd’, probably at MID, isotypes US964504, GH58549). UC1491 was collected 16 July 1897 and is apparently not type material.
PARATYPES: Yosemite Valley, H.N. Bolander 6196 (paratypes GH, DS, isoparatype UC930); Plumas County, Mrs. R.M. Austin (GH); Siskiyou County, Sisson, H.E. Brown 379 (NY). NOTE: WILLU24762 is listed as an isotype, but the specimen was not mentioned in the protologue of C. brainerdii.228. Carex brevipes W. Boott in S. Watson
Bot. Calif. 2:246. 1880.
TYPE: California, Sierra Nevada, Lake Tahoe to Bear Valley, A. Kellogg (possible types at GH57538 and NY25133), NY specimen shows the date of 9 June 1870 and is attributed to Brainerd & Kellogg.
NOTE: in California, this taxon seems quite distinct from C. rossii in which it is often submerged, or at least there are two taxa in this complex, regardless of applied name. The UC/JEPS specimens are on loan and not yet in the database.Carex rossii var. brevipes Kuk.
Pflanzenriech 4 (Heft 20):452. 1909.229. Carex rossii F. Boott in Hook. | Jepson Manual Distribution | Specimens |
Fl. Boreali Amer. 2:222. 1839.
SYNTYPES: northwest coast North America, D. Douglas; Rocky Mountains, Thomas Drummond (GH58557).
DISTRIBUTIONAL NOTE: the stated Jepson Manual range omits reference to New Mexico stations (Roalson, E.H., Jones, S.D., Allred, K.W. Sida 16(3): 592-594. 1995)232. Carex brevicaulis Mack. | Jepson Manual Distribution | Specimens |
Bull. Torrey Bot. Club. 40:547. 1913.
TYPE: Oregon, Yaquina Bay, T.J. Howell 2994 (type NY21532)
PARATYPES: British Columbia, Victoria, J.M. Macoun 76706, June 12, 1908 (NY); Vancouver, J.M.Macoun, May 30, 1873 (GH); J.M. Macoun 32014, May 8, 1875 ("D.C."); Washington, Whidbey Island, Gardner 343, May 29, 1897 (Piper); Wilkes Exped. 1834-1842 (C, = NY?); California, San Francisco, A. Kellogg, May 1880 (NY).225a. Carex serpenticola P.F. Zika29. Sect. Digitatae Fries (Fries) Christ
Madrono 45(3):261. 1998 (published 22 June 1999)
TYPE: California, Del Norte County, SE of Azalea Lane, N of Middle Fork Smith river, ca. 0.8 km SE of bridge in the town of Gasquet, 41°511N, 123°57.5'W (T17N R2E Section 21, SW1/4, Humboldt Meridian), elev. 120 m., 29 April 1997, P.F.Zika and K. Kuykendall 13062 (Holotype OSC, isotypes HSC, MICH, NY, UC, US).
(the types are as yet not online 12/12/00)
NOTE: a special status plant in California (CNPS List 2, vouchers as above), and Oregon.
Fl. Scan. 187. 1835. & Bull. Bot. Soc. Belg. 32:13. 1885
SYNOPSIS: dry ground calciphiles of continental and temperate Eurasia and North America, where Mackenzie recognized 4 species. Not known from South America, Australasia nor South Africa. TYPE: Carex digitata L. (Europe)245. Carex concinnoides Mack. | Jepson Manual Distribution | Specimens |38. Sect. Bicolores Tuck.
Bull. Torrey Bot. Club. 33:439. 1906.
TYPE: Montana, Columbia Falls, Williams in 1893.
Enum. Caric. 12. 1843.
SYNOPSIS: 5 calciphile or halophiles of N Hemisphere, south in the mountains of western North America. Egorova would (perhaps correctly) place our species in Sect. Paniceae. TYPECarex bicolor All.271. Carex hassei L.H.Bailey | Jepson Manual Distribution | Specimens |
Bot. Gaz. 21:5. 1896.
TYPE: California, San Gabriel Mountains, San Antionio Canyon, H.E. Hasse s.n., June 1884 (holotype BH?, isotypes UC115918, NY11140, NY11141). Mackenzie listed July as the collection date, but the protologue reads June as does the UC sheet (the NY sheets have a July date - the discrepancy may point to the NY material as being non-types).
DISTRIBUTIONAL NOTE: the stated Jepson Manual range does not reflect a single n Baja CA station in the Sierra de San Pedro Martir.Carex aurea var. celsa L.H. Bailey
Mem. Torrey Bot. Club 1:75. 1889.
SYNTYPES: California, San Bernardino Mountains, G.R. Vasey s.n., Portland Oregon, L.F. Henderson, T. Howell. Cronquist says about this "the robust extreme" (Vasc. Plants Pacific NW)Carex celsa Piper in Piper & Beattie
Fl. N.W. Coast 79. 1915.
TYPE: based on C. aurea var. celsa L.H. Bailey?270. Carex saliniformis Mack. | Jepson Manual Distribution | Specimens |
Bull. Torrey Bot. Club 36:477. 1909.
TYPE: based on C. salina var. minor W. Boott.
Original spelling "salinaeformis" changed to conform to ICBN Article 60.8
NOTE: a special status plant endemic to California (CNPS List 1B, vouchered from Humboldt, Mendocino, Sonoma and Santa Cruz [extirpated] counties)Carex salina var. minor W. Boott in S. Watson
Bot. Calif. 2:242. 1880.
TYPE: California, Mendocino Co., swamps near Mendocino, H.N. Bolander 4702 (possible types GH56313 & GH56314, isotypes, UC1278, UC1277, UC1279, JEPS25941, CAS383801, US29888 & US319226).272. Carex aurea Nutt.|Jepson Manual Distribution | Specimens |
Gen. N. Amer. Pl. 2:205. 1818.
TYPE: shores of Lake Michigan, Nuttall (holotype K?).
NOTE: Placed in Section Paniceae (Carey) Christ by Egorova272a. Carex garberi Fern. | Jepson Manual Distribution| Specimens |39. Sect. Paniceae Tuck.
Rhodora 37:253. 1935.
TYPE: Pennsylvania, Erie Co., Presque Isle, Garber s.n., June 9, 1869 (holotype GH?, isotypes NY32680,
US63525,NY32678, NY32675)
NOTE: Placed in Section Phacocystis, Subsection Cynaecandrae Egor. by Egorova
Enum. Caric. 15. 1943.
SYNOPSIS: a group of about 12 species, in dry to swamps, in cool temperate North America and Eurasia. Mackenzie says two South American species have been referred to the Paniceae. TYPE C. panicea L.273. Carex livida (Wahlenb.) Willd.
Sp. Pl. 4:285. 1805. | Jepson Manual Distribution| Specimens |
TYPE: based on C. limosa var. livida Wahl.
NOTE: A special status plant in California (CNPS List 1A, presumed extirpated) Oregon, Idaho. H.N. Bolander 4745 (UC8042), from Mendocino is the only CA station.Carex limosa var. livida Wahl.281. Carex californica L.H.Bailey
Sv. Vet.-Akd. Nya. Handl. 24: 162. 1803
TYPE: from Lapland
Mem. Torrey Bot. Club. 1:9. 1889. | Jepson Manual Distribution| Specimens |
TYPE: California, Mendocino County, Mendocino, H.N. Bolander 4741, 1 May 1866 (isotypes US29741),US319268, DS49734). UC1207 & UC689 are isotypes, but these specimens are not in the UCJEPS accessions database.
NOTE: this species is nowhere common, but geographically widespread. It is a special status plant in California (CNPS List 2, where vouchered only from Mendocino County), Idaho.40. Sect. Laxiflorae Kunth.Carex polymorpha var. californica (L.H. Bailey) Kuk.
Das Pflanzenreich 4, 20 (Heft 38):515. 1909.
TYPE: based on C. californica
Enum. Pl. 2:452 in part. 1837.
SYNOPSIS: a section best developed in the forests of Eastern North America; one species in the western U.S., and several species in eastern Asia. (cf. Anomalae). TYPE: Carex laxiflora L.293. Carex hendersonii L.H.Bailey44. Sect. Longicules Mack.
Proc. Amer. Acad. 22: 115. 1886. | Jepson Manual Distribution| Specimens |
LECTOTYPE: bogs at Portland, L. F. Henderson (the ORE95848 sheet is listed as an isosyntype, and was collected 20 May 1882, GH27267 is listed as a syntype, NY11147, NY11146, OSC1929 is listed as an isolectotype; the two ORE/OSC specimens were collected on different dates. The lectotype by Mackenzie op. cit. p. 253, 1935, but neither of the NY specimens is clearly labeled as the lectotype.
SYNTYPES: "Lower Frazier River, lat. 49°, Dr. Lyall; Oregon, (E. Hall 602); Multnomah Co., T. Howell [UC115920 & UC1184897 are a probable syntypes], and probably Nuttall’s specimens cited by Boott (Ill. 36; Mendocino County, Calif, Bolander 4747)" [the latter therefore paratypes, see UC1040 and UC963].
Fl. North Amer. 18:271. 1931.
SYNOPSIS: a group of 5 dry ground species, 1 in California, 4 in N Mexico (i.e. Madrean) 1 in Guatemala (C. guatemalensis). Carex triquerta shows some affinities to this section and may requirement relocation hereto.321. Carex whitneyi Olney | Jepson Manual Distribution| Specimens |
Proc. Amer. Acad. 7:394. 1868.
LECTOTYPE: California, Yosemite Valley, W.H. Brewer 1639 [lectotype NY11413]. Lectotype designated by Mackenzie (Erythea 8:61. 1922.)
SYNTYPES: Bolander 6198 [NY11412], Hillebrand 2305, 2308, 2314; Mt. Dana, 12000 ft, Bolander 5086 [NY11411, NY14110, UC1525, UC1530, UC70621]; Soda Springs, 9000 ft, Brewer 1778 [NY11409, UC1531, UC8007]". The three UC sheets of Bolander 5086 each bears different locality information, UC1530 from 12000 ft on Mt. Dana which is ecologically erroneous for this taxon. The CAS type catalogue lists an isotype in their collection (DS54663), but this specimen in a syntype. UC1049 is a sheet of Bolander 6198 that is Carex lenticularis var. impressa, and is therefore not a syntype of C. whitneyi, but is rather a paratype of C. paucicostata.Carex flaccifolia Mack.
Erythea 8:92. 1922.
TYPE: 'southwestern California', George B. Grant, 1 May 1902 (US468192). Howell (Leaf. W. Bot. 8: 221-224. 1958) concluded that the specimen was actually collected in Yosemite Valley.47b Sect. Silvaticae Rouy. Fl. France 13:1912Carex jepsonii J. Howell
Leaf. W. Bot. 8:223. 1958.
TYPE: California, Tuolumne County, Tuolumne Meadows, W.L.Jepson 4477 (holotype NY11183, no material at JEPS).
(Sect. Sylvaticae F. Boott in Tuck. Enum. Caric. 12. 1843. of Mackenzie 1935)
SYNOPSIS: a polyphylletic group of meadow and wetland species, with about 40 species. Work by Waterway & Olmstead (unpub.) suggests our species formerly placed in Sect. Sylvaticae and a core group of Sect. Gracillimae Carey form a monophylletic group related to Sect. Hymenochlaneae Drej. ex L.H. Bailey. There are perhaps two undescribed California taxa in this section (cf. L. Janeway 1992. Cyperaceae of Butte County, Cont. Herb. Chico State Univ.), both in the Sierra Nevada, requiring study.333. Carex gynodynama Olney
Proc. Amer. Acad. 7:394. 1868.| Jepson Manual Distribution| Specimens |
TYPE: California, Mendocino County, near Mendocino, H.N. Bolander 4700 (2 types at US28978 & US319267, isotype DS49500 ). No Bolander material of this at UC has been found.
NOTE: a special status plant in Oregon.Carex blankinshipii Fernald
Erythea 7:121. 1899.
TYPE: California, Humboldt County, Hydesville, J.W. Blankinship, June 19, 1893 (holotype GH?). The type does not appear in the GH types database.334. Carex mendocinensis Olney ex S. Watson
Bot. Calif. 2:249. 1880.| Jepson Manual Distribution| Specimens |
TYPE: in swamps near Mendocino, H.N. Bolander 4701 (isotypes US29453, DS54832). No Bolander material of this at UC. The plants in the Sierra Nevada may represent an undescribed taxon.Carex debiliformis Mack.
Bull. Torrey Bot. Club 37:244. 1910.
TYPE: based on C. cinnamoena Olney, not C. cinnamomea Boott, 1846.Carex cinnamoena Olney in A. Gray
Proc. Amer. Acad. 8:396. 1872.
TYPE: California, Mendocino County, H.N. Bolander 6477. (type fragments US28457, US319228 from ‘Red Mountains’ isotype CAS553874). No Bolander material at UC.335. Carex hirtissima W. Boott in S. Watson48. Sect. Cryptostachyae Meinsh.
Bot. Calif. 2:247. 1880.-|Jepson Manual Distribution| Specimens |
TYPE: in the Sierra Nevada, A. Kellogg (holotype not listed in GH catalog, nor did Waterway [Madrono 43:1-14. 1996 report on its status or whereabouts]335a. Carex obispoensis Stacey
Leaf. W. Bot. 1:240. 1936.| Jepson Manual Distribution| Specimens |
TYPE: California, San Luis Obispo County, Steiner Creek near San Luis Obispo, A. Eastwood & J.T. Howell 2271, May 7, 1936 (holotype CAS253733, isotypes CAS237908, US1678188, WTU77900). Paratype: I.J. Condit, April 1910 (cited as UC, but not found).
NOTE: a special status plant endemic to California (CNPS List 1B, vouchered only San Luis Obispo county and reported from Monterey County, but not vouchered therefrom)
