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Vascular Plants of California
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Myriopteris covillei
COVILLE'S LIP FERN


Higher Taxonomy
Family: PteridaceaeView DescriptionDichotomous Key
Common Name: BRAKE FAMILY
Habit: Perennial herb, in soil or on or among rocks; rhizome creeping to erect, scaly. Leaf: generally all +- alike (or of 2 kinds, fertile, sterile), generally < 50 cm, often < 25 cm; stipe generally thin, wiry, often dark, ×-section with vascular strands generally 1--3, less often many in circle; blade generally pinnate or +- palmate-pinnate (see Adiantum), often >= 2-compound, abaxially often with glands, +- powdery exudate, hairs, or scales; segments round, oblong, fan-shaped, or other, veins generally free. Sporangia: in sori or not, marginal, submarginal, or along veins, covered by recurved, often modified segment margins (false indusia) or not; true indusia 0; spores spheric, sides flat or not, scar with 3 radiating branches.
Genera In Family: +- 40 genera, 500 species: worldwide, especially dry areas. Note: CA members of Cheilanthes moved to the distantly related Myriopteris; Pellaea breweri to be moved as well, from a to-be-redefined Pellaea; traditional, often untenable limits of genera outside CA also being clarified using molecular phylogenetics.
eFlora Treatment Author: Ruth E.B. Kirkpatrick, Alan R. Smith & Thomas Lemieux, except as noted
Scientific Editor: Alan R. Smith, Thomas J. Rosatti.
Genus: MyriopterisView DescriptionDichotomous Key


Habit: Plant in soil or rock crevices; rhizome short- to long-creeping[-decumbent], generally many-branched, scales generally lance-linear, pale to dark, mid-stripe dark or not. Leaf: < 75 cm, young leaf tip hooked or coiled; stipe cylindric, red-brown to +- black; blade generally 2--3-pinnate, generally oblong to narrowly triangular; segments generally small, +- flat or abaxially concave (from recurved margins). Sporangia: along margin, in discrete patches to continuous, partly to completely covered by recurved margin (generally not recurved in Myriopteris cooperae).
Etymology: (Greek: myriad fern, from much divided leaf blades) Note: Includes all CA members of Cheilanthes as treated in TJM2 (2012).
eFlora Treatment Author: Ruth E.B. Kirkpatrick, Alan R. Smith, Thomas Lemieux & Edward Alverson
Reference: Grusz & Windham 2013 PhytoKeys 32: 49--64, doi: 10.3897/phytokeys.32.6733
Myriopteris covillei (Maxon) Á. Löve & D. Löve
NATIVE
Habit: Rhizome short-creeping; scales dark brown or black, some with narrow light brown margin. Leaf: 8--22(30+) cm, 2--4(6) cm wide, dark green; stipe < 2 mm wide, scales lance-linear, +- white to red-brown; blade 3--4-pinnate; segments small, +- round, adaxially glabrous, abaxially concave, obscured by scales exceeding margin, scales originating from axes, > 2 mm, +- > 1 mm wide, +- entire, covering generally more highly dissected scales. Sporangia: obscured by recurved segment margin, scales. Chromosomes: 2n=60.
Ecology: Crevices, bases of rocks, sun or shade; Elevation: 600--2400 m. Bioregional Distribution: NCoR, SN, ScV (Sutter Buttes), SnFrB, SCoR, e SCo, TR, PR, SNE, w edge DMoj, DMtns, w DSon; Distribution Outside California: to Utah, Arizona, Baja California. Note: Hybrids, presumed sterile, occasionally with Myriopteris intertexta, Myriopteris newberryi (Myriopteris ×fibrillosa (Davenp.) Grusz & Windham), Myriopteris parryi (Myriopteris ×parishii (Davenp.) Grusz & Windham) in southern California.
Synonyms: Cheilanthes covillei Maxon
Jepson eFlora Author: Ruth E.B. Kirkpatrick, Alan R. Smith, Thomas Lemieux & Edward Alverson
Reference: Grusz & Windham 2013 PhytoKeys 32: 49--64, doi: 10.3897/phytokeys.32.6733
Index of California Plant Names (ICPN; linked via the Jepson Online Interchange)

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Botanical illustration including Myriopteris covillei

botanical illustration including Myriopteris covillei

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Citation for this treatment: Ruth E.B. Kirkpatrick, Alan R. Smith, Thomas Lemieux & Edward Alverson 2014, Myriopteris covillei, in Jepson Flora Project (eds.) Jepson eFlora, Revision 2, https://ucjeps.berkeley.edu/eflora/eflora_display.php?tid=99420, accessed on April 19, 2024.

Citation for the whole project: Jepson Flora Project (eds.) 2024, Jepson eFlora, https://ucjeps.berkeley.edu/eflora/, accessed on April 19, 2024.

Myriopteris covillei
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©2024 Neal Kramer
Myriopteris covillei
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©2015 Barry Breckling
Myriopteris covillei
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©2023 Dylan Neubauer
Myriopteris covillei
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©2024 Neal Kramer
Myriopteris covillei
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©2024 Neal Kramer

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Geographic subdivisions for Myriopteris covillei:
NCoR, SN, ScV (Sutter Buttes), SnFrB, SCoR, e SCo, TR, PR, SNE, w edge DMoj, DMtns, w DSon
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).





 

Data provided by the participants of the  Consortium of California Herbaria.
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Blue markers indicate specimens that map to one of the expected Jepson geographic subdivisions (see left map). Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa, if there are more than 1 infraspecific taxon in CA.
Blue line denotes eFlora flowering time (fruiting time in some monocot genera).