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Vascular Plants of California
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Pellaea breweri
BREWER'S CLIFF-BRAKE


Higher Taxonomy
Family: PteridaceaeView DescriptionDichotomous Key
Common Name: BRAKE FAMILY
Habit: Perennial herb, in soil or on or among rocks; rhizome creeping to erect, scaly. Leaf: generally all +- alike (or of 2 kinds, fertile, sterile), generally < 50 cm, often < 25 cm; stipe generally thin, wiry, often dark, ×-section with vascular strands generally 1--3, less often many in circle; blade generally pinnate or +- palmate-pinnate (see Adiantum), often >= 2-compound, abaxially often with glands, +- powdery exudate, hairs, or scales; segments round, oblong, fan-shaped, or other, veins generally free. Sporangia: in sori or not, marginal, submarginal, or along veins, covered by recurved, often modified segment margins (false indusia) or not; true indusia 0; spores spheric, sides flat or not, scar with 3 radiating branches.
Genera In Family: +- 40 genera, 500 species: worldwide, especially dry areas. Note: CA members of Cheilanthes moved to the distantly related Myriopteris; Pellaea breweri to be moved as well, from a to-be-redefined Pellaea; traditional, often untenable limits of genera outside CA also being clarified using molecular phylogenetics.
eFlora Treatment Author: Ruth E.B. Kirkpatrick, Alan R. Smith & Thomas Lemieux, except as noted
Scientific Editor: Alan R. Smith, Thomas J. Rosatti.
Genus: PellaeaView DescriptionDichotomous Key


Common Name: CLIFF-BRAKE
Habit: Plant in soil or rock crevices; rhizome short- to long-creeping, scales overlapped, narrowly linear, light- to red- or medium-brown, often with dark mid-stripe. Leaf: erect, persistent, < 1 m; stipes +- cylindric, generally dark or red-brown to +- black, +- shiny, glabrous; blade 1--4-pinnate; segments generally stalked, generally free, linear to rounded, lobed or not, often folded lengthwise when dried; veins generally free. Sporangia: in +- continuous, submarginal bands, among a +- white to +- yellow exudate or not; segment margin generally recurved, generally modified; spores tan to light yellow.
Etymology: (Greek: dusky, from blue-gray leaves) Note: Occasionally cultivated. Molecular data suggest Pellaea in the sense of Tryon (1957) is polyphyletic with monophyletic Astrolepis and Pellaea sect. Platyloma, as well as elements of Paragymnopteris and Paraceterach nested within (Kirkpatrick, 2007).
eFlora Treatment Author: Ruth E.B. Kirkpatrick, Alan R. Smith, Thomas Lemieux & Edward Alverson
Reference: Kirkpatrick 2007 Syst Bot 32:504--518
Unabridged Reference: Kirkpatrick, R.E.B. 2007. Investigating the monophyly of Pellaea (Pteridaceae) in the context of a phylogenetic analysis of cheilanthoid ferns. Syst Bot 32:504--518; Tryon 1957 Ann Missouri Bot Gard 44(2):125--193
Pellaea breweri D.C. Eaton
NATIVE
Habit: Rhizome short-creeping, branched, > 10 cm, 5(7) mm wide; scales +- thread-like, red-brown, dark mid-stripe 0. Leaf: clustered, 8--20(25) cm, 2--3(4) cm wide, pale greenish; stipe < 2 mm wide, fracture lines at base many; blade 1-pinnate, oblong, rachis green at tip; pinnae < 2 cm, < 1.5 cm wide, lance-ovate, deeply 2(3)-lobed. Sporangia: 64-spored; spores dark to light brown. Chromosomes: 2n=58.
Ecology: Generally n-facing granite rock crevices, slopes; Elevation: 1500--3700 m. Bioregional Distribution: KR, CaRH, SNH, SnBr, GB, DMtns; Distribution Outside California: to Washington, Idaho, Colorado. Note: Molecular evidence suggests that Paragymnopteris marantae (L.) K.H. Shing of Eurasia and Paragymnopteris sargentii (H. Christ) K.H. Shing of China are sister to each other, that both are sister to Pellaea breweri, and that these 3 form the earliest diverging pellaeoid clade (Kirkpatrick 2007).
Jepson eFlora Author: Ruth E.B. Kirkpatrick, Alan R. Smith, Thomas Lemieux & Edward Alverson
Reference: Kirkpatrick 2007 Syst Bot 32:504--518
Index of California Plant Names (ICPN; linked via the Jepson Online Interchange)

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Botanical illustration including Pellaea breweri

botanical illustration including Pellaea breweri

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Citation for this treatment: Ruth E.B. Kirkpatrick, Alan R. Smith, Thomas Lemieux & Edward Alverson 2012, Pellaea breweri, in Jepson Flora Project (eds.) Jepson eFlora, https://ucjeps.berkeley.edu/eflora/eflora_display.php?tid=36778, accessed on April 16, 2024.

Citation for the whole project: Jepson Flora Project (eds.) 2024, Jepson eFlora, https://ucjeps.berkeley.edu/eflora/, accessed on April 16, 2024.

Pellaea breweri
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©2004 Larry Blakely
Pellaea breweri
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©2011 Barry Breckling
Pellaea breweri
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©2004 Larry Blakely
Pellaea breweri
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©1998 Larry Blakely
Pellaea breweri
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©2005 Dieter Wilken

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Geographic subdivisions for Pellaea breweri:
KR, CaRH, SNH, SnBr, GB, DMtns
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).





 

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Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa, if there are more than 1 infraspecific taxon in CA.
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