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Vascular Plants of California
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Carduus acanthoides subsp. acanthoides

Higher Taxonomy
Family: Asteraceae (Compositae)View DescriptionDichotomous Key
Habit: Annual to tree. Leaf: basal and/or cauline, alternate, opposite, rarely whorled, simple to 2+ × compound. Inflorescence: 1° inflorescence a head, resembling a flower, of several types (see below), 1--many in generally +- cyme-like cluster; each head generally with +- calyx-like involucre of 1--many series of phyllaries (involucre bracts); receptacle of head flat to conic or columnar, paleate (bearing paleae = receptacle bracts) or epaleate; flowers 1--many per head. Flower: bisexual, unisexual, or sterile, +- small, of several types (see below); calyx 0 or modified into +- persistent pappus of bristles, scales, and/or awns; corolla radial or bilateral (0), lobes generally (0)3--5; stamens 4--5, filaments generally free, generally fused to corolla at tube/throat junction, anthers generally fused into cylinder around style, anther base generally rounded or cordate (deeply sagittate or with tail-like appendages), tip (= flattened appendage) generally projecting beyond pollen sac; pistil 1, 2-carpeled, ovary inferior, 1-chambered, 1-seeded, placenta basal, style 1, tip generally +- 2-branched (except in some staminate disk flowers), branch tips truncate or generally bearing +- brush-like appendages; stigmas 2, generally on adaxial faces of style branches. Fruit: achene (also called a cypsela) (drupe in Chrysanthemoides), cylindric to ovoid, sometimes compressed, generally deciduous with pappus attached.
Genera In Family: +- 1500 genera, 23000 species: worldwide, many habitats. Note: Flower and head types differ in form and sexual condition. A disk flower has a generally radial corolla, with a cylindric tube, expanded throat, and generally 5 lobes. Disk flowers are generally bisexual and fertile but occasionally staminate with reduced ovaries. Discoid heads comprise only disk flowers. A radiant head is a variant of a discoid head, with peripheral disk flower corollas expanded, often bilateral. A ray flower corolla is bilateral, generally with a slender tube and flattened petal-like ray (single lip composed of generally 3 lobes). Ray flowers are generally pistillate or sterile (occasionally lacking styles). Radiate heads have peripheral ray flowers and central disk flowers. Disciform heads superficially resemble discoid heads, with pistillate or sterile flowers that lack rays, together with or separate from disk flowers. A ligulate flower is bisexual, with a bilateral, generally ephemeral corolla and 5-lobed ligule. Liguliflorous heads comprise only ligulate flowers. See glossary p. 31 for illustrations of family characteristics. Echinops sphaerocephalus L., Gaillardia aristata Pursh, Gaillardia pulchella Foug., Hymenothrix loomisii S.F. Blake, Tagetes erecta L., Thelesperma megapotamicum (Spreng.) Kuntze are waifs. Melampodium perfoliatum Kunth, historic urban waif. Ageratum conyzoides L., Guizotia abyssinica (L. f.) Cass., Santolina chamaecyparisus L., orth. var. are rare or uncommon escapes from cultivation. Dyssodia papposa, Ismelia carinata (Schousb.) Sch. Bip. [Chrysanthemum carinatum Schousb.], Mantisalca salmantica (L.) Briq. & Cavill. are historical or extirpated waifs in California. Inula helenium L. not documented in California. Taxa of Aster in TJM (1993) treated here in Almutaster, Eucephalus, Eurybia, Ionactis, Oreostemma, Sericocarpus, Symphyotrichum; Chamomilla in Matricaria; Bahia in Hymenothrix; Cnicus in Centaurea; Conyza in Erigeron and Laennecia; Dugaldia in Hymenoxys; Erechtites in Senecio; Hymenoclea in Ambrosia; Lembertia in Monolopia; Osteospermum ecklonis in Dimorphotheca; Picris echioides in Helminthotheca; Prionopsis in Grindelia; Raillardiopsis in Anisocarpus and Carlquistia; Schkuhria multiflora in Picradeniopsis; Trimorpha in Erigeron; Venidium in Arctotis; Whitneya in Arnica. Amauriopsis in TJM2 (2012) treated here in Hymenothrix; Arida in Leucosyris; Bahia in Picradeniopsis.
Unabridged Note: Largest family of vascular plants in California and of eudicots globally.
eFlora Treatment Author: David J. Keil, except as noted
Scientific Editor: David J. Keil, Bruce G. Baldwin.
Genus: CarduusView DescriptionDichotomous Key

Habit: Annual, biennial [perennial herb]. Stem: erect. Leaf: simple, basal and cauline, alternate, reduced upward, decurrent as spiny wings, spiny-dentate and pinnately lobed, glabrous to tomentose; basal tapered to winged petiole; cauline sessile. Inflorescence: heads discoid, 1--20 at branch tips; involucre cylindric to spheric; phyllaries graduated in several series, spine-tipped; receptacle flat, epaleate, +- white bristly. Flower: corolla white to pink or purple, tube long, slender, throat abruptly expanded, short, lobes linear; anther base short-sagittate, tip oblong; style tip cylindric above slightly swollen distal node, branches very short. Fruit: ovoid, slightly compressed, glabrous; base slightly angled; pappus of many flat, minutely barbed bristles.
Species In Genus: +- 90 species: Eurasia, eastern Africa. Etymology: (Latin: ancient name for a kind of thistle)
eFlora Treatment Author: David J. Keil
Reference: Keil 2006 FNANM 19:91--94
Carduus acanthoides L. subsp. acanthoides
Habit: Biennial. Stem: 3--15 dm, +- glabrous to loosely woolly, strongly spiny-winged. Leaf: basal 10--30 cm, spiny-toothed to deeply 1--2-pinnately lobed, generally sparsely hairy. Inflorescence: heads 1--5 in clusters; involucre 1--2.5 cm diam, +- spheric or hemispheric; phyllary base glabrous or thinly tomentose, margin not scarious, tip linear or narrowly lanceolate, appressed to spreading, minutely scabrous-ciliate. Flower: corolla 13--20 mm, purple; tube 5--10 mm, throat +- 2 mm, lobes 6--8 mm. Fruit: 2.5--3 mm, golden to brown; pappus 11--13 mm. Chromosomes: 2n=22.
Ecology: Roadsides, pastures, disturbed areas; Elevation: < 1300 m. Bioregional Distribution: NCoRI, n SN, n CCo, n SCoRO, MP; Distribution Outside California: North America; native to Europe. Flowering Time: Jul--Nov
Jepson eFlora Author: David J. Keil
Reference: Keil 2006 FNANM 19:91--94
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Botanical illustration including Carduus acanthoides subsp. acanthoides

botanical illustration including Carduus acanthoides subsp. acanthoides


Citation for this treatment: David J. Keil 2012, Carduus acanthoides subsp. acanthoides, in Jepson Flora Project (eds.) Jepson eFlora,, accessed on December 16, 2018.

Citation for the whole project: Jepson Flora Project (eds.) 2018, Jepson eFlora,, accessed on December 16, 2018.

No expert verified images found for Carduus acanthoides subsp. acanthoides.

Geographic subdivisions for Carduus acanthoides subsp. acanthoides:
NCoRI, n SN, n CCo, n SCoRO, MP;
Markers link to CCH specimen records. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues. Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa, if there are more than 1 infraspecific taxon in CA.
Blue line denotes eFlora flowering time.