Common Name: HEATH FAMILY
Habit: Perennial herb, shrub, tree. Stem: bark often peeling distinctively. Leaf: simple or 0, generally cauline, alternate, opposite (whorled), evergreen or deciduous, often leathery, petioled or not; stipules 0. Inflorescence: raceme, panicle, cyme, or flowers 1, terminal or axillary, generally bracted; pedicel often with 2 bractlets. Flower: generally bisexual, generally radial, bell-shaped, cylindric, or urn-shaped; sepals generally (0)4--5, generally free; petals generally (0)4--5, free or fused; stamens (2--5)8--10, free, filaments rarely appendaged, anthers dehiscing by pores or slits, awns 0 or 2(4), seemingly abaxial, reduced or elongate, generally curved; nectary generally present at ovary base, generally disk-like; ovary superior or inferior, chambers generally 1--5, placentas axile or parietal, ovules 1--many per chamber, style 1, stigma head- to funnel-like or lobed. Fruit: capsule, drupe, berry. Seed: generally many, winged or not.
Genera In Family: +- 100 genera, 3000 species: generally worldwide except deserts; some cultivated, especially Arbutus, Arctostaphylos, Rhododendron, Vaccinium. Note: Monophyletic only if Empetraceae included, as treated here. Ledum included in Rhododendron. Non-green plants obtain nutrition from green plants through fungal intermediates.
eFlora Treatment Author: Gary D. Wallace, except as noted
Scientific Editor: Gary D. Wallace, Thomas J. Rosatti.
Common Name: MANZANITA
Habit: Shrub to small tree, prostrate to erect. Stem: old stems generally +- red, smooth, bark generally thin, peeling, or generally +- gray or red-gray, shredding and rough; burls at base, woody, sprouting after fire, or generally 0; twig hairs 0 or generally +- like those on inflorescence axes, bracts. Leaf: alternate, evergreen; blade flat to convex, base lobed to wedge-shaped, clasping stem or not, margins generally flat, surfaces with stomata generally both abaxially, adaxially, alike in color, hairiness, less often only or fewer abaxially, generally differing in color, hairiness. Inflorescence: +- raceme (generally 0--1-branched) or panicle (generally 2--10-branched), terminal, nascent inflorescence present following stem growth, generally late spring through winter, remaining dormant 4--6 months prior to flower (except in Arctostaphylos pringlei subsp. drupacea); branches 0 or raceme-like; flower bracts leaf-like, generally flat, or scale-like, often folded, keeled, tips rounded to acute to awl-shaped. Flower: radial; sepals 5(4), free, persistent; corolla conic to urn-shaped, lobes in number = sepals, short, rounded, curved back, white to pink; stamens 2 × number of sepals, included, filaments swollen, generally hairy at base, anthers dark red, awns elongate; ovary superior, on disk, 4--10-chambered, ovule 1 per chamber. Fruit: drupe, generally +- depressed-spheric to spheric; flesh generally thick, +- mealy, occasionally 0; stones 2--10, free, fused, or some fused.
Species In Genus: +- 62 species: North America (especially California) to Central America, Eurasia. Etymology: (Greek: bear berries)
Unabridged Note: Rosatti (1986 Syst Bot 12:61--77) showed that in Arctostaphylos uva-ursi little to none of the variation in hairs (including length, glandularity) is genetically based.
eFlora Treatment Author: V. Thomas Parker, Michael C. Vasey & Jon E. Keeley
Habit: Erect, 1--3 m; burl prominent. Stem: twig glabrous or tomentose, glandular or not. Leaf: spreading; petiole 2--5 mm; blade 2--5 cm, 1.5--2.5 cm wide, oblong-ovate to lance-oblong, abaxially glabrous to densely short-nonglandular-hairy, light green, adaxially +- glabrous (nonglandular-hairy), dark green, base truncate to +- lobed, tip acute, margin entire, occasionally toothed, cupped or +- rolled; stomata abaxial. Inflorescence: panicle, 2--8-branched; nascent inflorescence pendent, axis 1--2.5 cm, > 1 mm wide, glabrous, nonglandular-hairy, or glandular-hairy; lower bracts leaf-like, upper often scale-like, tomentose, glandular or not; bracts 3--8 mm, scale-like and awl-shaped to deltate or leaf-like and lanceolate, glabrous to hairy; pedicel 2--5 mm, sparsely nonglandular-hairy to glandular-hairy. Flower: ovary glabrous to short-hairy, generally nonglandular. Fruit: 6--10 mm wide, depressed-spheric, glabrous to sparsely nonglandular-hairy; stones free. Chromosomes: 2n=52.