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Nemacladus secundiflorus var. secundiflorus

Higher Taxonomy
Family: CampanulaceaeView DescriptionDichotomous Key

Habit: Annual to perennial herb [tree]. Leaf: generally cauline, generally simple, generally alternate, petioled or not; stipules 0. Inflorescence: cyme, panicle, raceme, spike, or flowers 1; terminal or in axils of leaf-like or reduced bracts. Flower: bisexual, cleistogamous or open, radial or bilateral, inverted (pedicel twisted 180°) or not; hypanthium generally present, +- fused to ovary; sepals generally 5; corolla radial to 2-lipped, petals generally fused, tube deeply divided on 1 side or not, lobes generally 5; stamens 5, free or +- fused (anthers, filaments fused into tube or filaments fused above middle); ovary inferior or 1/2 inferior (superior in fruit), chambers 1--3, placentas axile or parietal, ovules many, style generally 1, 2--5-branched. Fruit: generally capsule, open on sides or top by pores or short valves. Seed: many.
Genera In Family: +- 90 genera, +- 2500 species: worldwide. Note: Some cultivated for ornamental (Campanula, Jasione, Lobelia). Subfamilies sometimes treated as families. Positions of flower parts given after flowering inversion, if any. Parishella moved to Nemacladus.
eFlora Treatment Author: Nancy R. Morin, except as noted
Scientific Editor: Thomas J. Rosatti.
Genus: NemacladusView DescriptionDichotomous Key

Habit: Annual, from taproot. Stem: prostrate, decumbent, or erect; base generally +- brown or +- purple; branches 0 or below middle. Leaf: basal; petiole short or 0. Inflorescence: +- raceme-like; bract 1 per flower, small; pedicel generally thread-like. Flower: inverted or not; sepals linear to triangular; corolla +- radial or 2-lipped, lobes 5; filaments free at base, fused into tube around style distally, appendages attached to a stalk or directly on 2 adjacent filaments, each with 2--12 cells, anthers free, all alike; ovary superior to 1/2 inferior, hemispheric to obconic, nectary glands 3, mounded or donut-like, on free part of ovary, stigma 2-lobed, papillate. Fruit: generally > hypanthium, hemispheric to fusiform, top pointed or rounded, chambers 2; open at top generally by 2 valves (or circumscissile). Seed: elliptic to oblong.
Species In Genus: 18 species: southwestern United States, northwestern Mexico. Etymology: (Greek: thread-like branch) Note: In descriptions, "filaments" including both free and fused parts thereof.
Unabridged Note: Taxonomic changes from TJM (1993) based on ITS, atpB, morphology. Parishella californica nested within Nemacladus, in which it is here treated.
Species: Nemacladus secundiflorusView Description 

Habit: Erect, 2.5--12.5(16) cm, branched from base or generally 2 cm above. Stem: spreading to ascending; base dull, red-brown. Leaf: 2--6 mm, narrowly oblanceolate to +- spoon-shaped or elliptic, irregularly dentate, hairy, narrowed to wide petiole. Inflorescence: +- 1-sided or not; axis straight or zigzag; bracts 1--3 mm, widely lanceolate to ovate or oblong, erect or reflexed; pedicels 8--15 mm, +- 0.1 mm diam, spreading, +- S-curved, tip erect. Flower: inverted; hypanthium 0.5 mm; sepals +- 0.8--2.5 mm, linear or deltate, erect; corolla salverform, divided 1/3 to base, lobes all held on lower side of flower; filaments arched, 0.8--2.5 mm, tip +- curved, long-hairy, appendages wide pads attached on filaments near tip of ovary, cells slender, pointed, anthers 0.1--0.7 mm; ovary +- 1/4-inferior. Fruit: +- 2--2.5 mm, base, tip rounded. Seed: 0.3--0.5 mm, +- spheric, with zigzag ridges alternate clearly pitted rows.

Nemacladus secundiflorus G.T. Robbins var. secundiflorus
Leaf: 3--6 mm, narrowly oblanceolate to +- spoon-shaped Inflorescence: +- 1-sided or not; axis straight; bracts 2--3 mm, widely lanceolate to oblong, erect or reflexed; pedicels 9--14 mm, +- 0.1 mm diam, horizontal, +- S-curved, tip erect. Flower: sepals +- 0.8--2.5 mm, linear; corolla white, with yellow blotch at lobe base and lavender veins, tube 2.5--3.5 mm, widely cylindric, 2 adaxial lobes spreading, 2--2.5 mm, glabrous to hairy, 3 abaxial reflexed, 2.5--3 mm; filaments +- 2--2.5 mm, anthers 0.5--0.7 mm.
Ecology: Dry, gravelly slopes; Elevation: 200--2000 m. Bioregional Distribution: s SNH, SCoR. Flowering Time: Apr--May
eFlora Treatment Author: Nancy R. Morin
Jepson Online Interchange
Listed on CNPS Rare Plant Inventory

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Citation for this treatment: Nancy R. Morin 2016. Nemacladus secundiflorus var. secundiflorus, in Jepson Flora Project (eds.) Jepson eFlora,, accessed on May 02, 2016.

Citation for the whole project: Jepson Flora Project (eds.) 2016. Jepson eFlora,, accessed on May 02, 2016.

Nemacladus secundiflorus var. secundiflorus
click for enlargement
© 2008 Aaron Schusteff
Nemacladus secundiflorus var. secundiflorus
click for enlargement
© 2008 Aaron Schusteff
Nemacladus secundiflorus var. secundiflorus
click for enlargement
© 2008 Aaron Schusteff

More photos of Nemacladus secundiflorus var. secundiflorus in CalPhotos

Geographic subdivisions for Nemacladus secundiflorus var. secundiflorus:
s SNH, SCoR.
Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.