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Nemacladus secundiflorus var. secundiflorus
LARGE-FLOWERED NEMACLADUS

Higher Taxonomy
Family: CampanulaceaeView DescriptionDichotomous Key

Common Name: BELLFLOWER FAMILY
Habit: Annual to perennial herb [tree]. Leaf: generally cauline, generally simple, generally alternate, petioled or not; stipules 0. Inflorescence: cyme, panicle, raceme, spike, or flowers 1; terminal or in axils of leaf-like or reduced bracts. Flower: bisexual, cleistogamous or open, radial or bilateral, inverted (pedicel twisted 180°) or not; hypanthium generally present, +- fused to ovary; sepals generally 5; corolla radial to 2-lipped, petals generally fused, tube deeply divided on 1 side or not, lobes generally 5; stamens 5, free or +- fused (anthers, filaments fused into tube or filaments fused above middle); ovary inferior or 1/2 inferior (superior in fruit), chambers 1--3, placentas axile or parietal, ovules many, style generally 1, 2--5-branched. Fruit: generally capsule, open on sides or top by pores or short valves. Seed: many.
Genera In Family: +- 90 genera, +- 2500 species: worldwide. Note: Some cultivated for ornamental (Campanula, Jasione, Lobelia). Subfamilies sometimes treated as families. Positions of flower parts given after flowering inversion, if any. Parishella moved to Nemacladus.
eFlora Treatment Author: Nancy R. Morin, except as noted
Scientific Editor: Thomas J. Rosatti.
Genus: NemacladusView DescriptionDichotomous Key

Habit: Annual, from taproot. Stem: prostrate, decumbent, or erect; base generally +- brown or +- purple; branches 0 or below middle. Leaf: basal; petiole short or 0. Inflorescence: +- raceme-like; bract 1 per flower, small; pedicel generally thread-like. Flower: inverted or not; sepals linear to triangular; corolla +- radial or 2-lipped, lobes 5; filaments free at base, fused into tube around style distally, appendages attached to a stalk or directly on 2 adjacent filaments, each with 2--12 cells, anthers free, all alike; ovary superior to 1/2 inferior, hemispheric to obconic, nectary glands 3, mounded or donut-like, on free part of ovary, stigma 2-lobed, papillate. Fruit: generally > hypanthium, hemispheric to fusiform, top pointed or rounded, chambers 2; open at top generally by 2 valves (or circumscissile). Seed: elliptic to oblong.
Species In Genus: 18 species: southwestern United States, northwestern Mexico. Etymology: (Greek: thread-like branch) Note: In descriptions, "filaments" including both free and fused parts thereof.
Unabridged Note: Taxonomic changes from TJM (1993) based on ITS, atpB, morphology. Parishella californica nested within Nemacladus, in which it is here treated.
Species: Nemacladus secundiflorusView Description 

Habit: Erect, 2.5--12.5(16) cm, branched from base or generally 2 cm above. Stem: spreading to ascending; base dull, red-brown. Leaf: 2--6 mm, narrowly oblanceolate to +- spoon-shaped or elliptic, irregularly dentate, hairy, narrowed to wide petiole. Inflorescence: +- 1-sided or not; axis straight or zigzag; bracts 1--3 mm, widely lanceolate to ovate or oblong, erect or reflexed; pedicels 8--15 mm, +- 0.1 mm diam, spreading, +- S-curved, tip erect. Flower: inverted; hypanthium 0.5 mm; sepals +- 0.8--2.5 mm, linear or deltate, erect; corolla salverform, divided 1/3 to base, lobes all held on lower side of flower; filaments arched, 0.8--2.5 mm, tip +- curved, long-hairy, appendages wide pads attached on filaments near tip of ovary, cells slender, pointed, anthers 0.1--0.7 mm; ovary +- 1/4-inferior. Fruit: +- 2--2.5 mm, base, tip rounded. Seed: 0.3--0.5 mm, +- spheric, with zigzag ridges alternate clearly pitted rows.


Nemacladus secundiflorus G.T. Robbins var. secundiflorus
NATIVE
Leaf: 3--6 mm, narrowly oblanceolate to +- spoon-shaped Inflorescence: +- 1-sided or not; axis straight; bracts 2--3 mm, widely lanceolate to oblong, erect or reflexed; pedicels 9--14 mm, +- 0.1 mm diam, horizontal, +- S-curved, tip erect. Flower: sepals +- 0.8--2.5 mm, linear; corolla white, with yellow blotch at lobe base and lavender veins, tube 2.5--3.5 mm, widely cylindric, 2 adaxial lobes spreading, 2--2.5 mm, glabrous to hairy, 3 abaxial reflexed, 2.5--3 mm; filaments +- 2--2.5 mm, anthers 0.5--0.7 mm.
Ecology: Dry, gravelly slopes; Elevation: 200--2000 m. Bioregional Distribution: s SNH, SCoR. Flowering Time: Apr--May
eFlora Treatment Author: Nancy R. Morin
Jepson Online Interchange
Listed on CNPS Rare Plant Inventory

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Citation for this treatment: Nancy R. Morin 2016. Nemacladus secundiflorus var. secundiflorus, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=89055, accessed on July 29, 2016.

Citation for the whole project: Jepson Flora Project (eds.) 2016. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on July 29, 2016.


Nemacladus secundiflorus var. secundiflorus
click for enlargement
© 2008 Aaron Schusteff
Nemacladus secundiflorus var. secundiflorus
click for enlargement
© 2008 Aaron Schusteff
Nemacladus secundiflorus var. secundiflorus
click for enlargement
© 2008 Aaron Schusteff

More photos of Nemacladus secundiflorus var. secundiflorus in CalPhotos


Geographic subdivisions for Nemacladus secundiflorus var. secundiflorus:
s SNH, SCoR.
Markers link to CCH specimen records. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues. Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.