Jepson eFlora: Taxon page
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Nemacladus californicus

Higher Taxonomy
Family: CampanulaceaeView DescriptionDichotomous Key

Habit: Annual to perennial herb [tree]. Leaf: generally cauline, generally simple, generally alternate, petioled or not; stipules 0. Inflorescence: cyme, panicle, raceme, spike, or flowers 1; terminal or in axils of leaf-like or reduced bracts. Flower: bisexual, cleistogamous or open, radial or bilateral, inverted (pedicel twisted 180°) or not; hypanthium generally present, +- fused to ovary; sepals generally 5; corolla radial to 2-lipped, petals generally fused, tube deeply divided on 1 side or not, lobes generally 5; stamens 5, free or +- fused (anthers, filaments fused into tube or filaments fused above middle); ovary inferior or 1/2 inferior (superior in fruit), chambers 1--3, placentas axile or parietal, ovules many, style generally 1, 2--5-branched. Fruit: generally capsule, open on sides or top by pores or short valves. Seed: many.
Genera In Family: +- 90 genera, +- 2500 species: worldwide. Note: Some cultivated for ornamental (Campanula, Jasione, Lobelia). Subfamilies sometimes treated as families. Positions of flower parts given after flowering inversion, if any. Parishella moved to Nemacladus.
eFlora Treatment Author: Nancy R. Morin, except as noted
Scientific Editor: Thomas J. Rosatti.
Genus: NemacladusView DescriptionDichotomous Key

Habit: Annual, from taproot. Stem: prostrate, decumbent, or erect; base generally +- brown or +- purple; branches 0 or below middle. Leaf: basal; petiole short or 0. Inflorescence: +- raceme-like; bract 1 per flower, small; pedicel generally thread-like. Flower: inverted or not; sepals linear to triangular; corolla +- radial or 2-lipped, lobes 5; filaments free at base, fused into tube around style distally, appendages attached to a stalk or directly on 2 adjacent filaments, each with 2--12 cells, anthers free, all alike; ovary superior to 1/2 inferior, hemispheric to obconic, nectary glands 3, mounded or donut-like, on free part of ovary, stigma 2-lobed, papillate. Fruit: generally > hypanthium, hemispheric to fusiform, top pointed or rounded, chambers 2; open at top generally by 2 valves (or circumscissile). Seed: elliptic to oblong.
Species In Genus: 18 species: southwestern United States, northwestern Mexico. Etymology: (Greek: thread-like branch) Note: In descriptions, "filaments" including both free and fused parts thereof.
Unabridged Note: Taxonomic changes from TJM (1993) based on ITS, atpB, morphology. Parishella californica nested within Nemacladus, in which it is here treated.

Nemacladus californicus (A. Gray) Morin
Habit: Prostrate, rosettes 1 or linked by naked, creeping stem, 1--10 cm diam. Stem: reclined. Leaf: 5--20 mm, oblanceolate, entire, hairy, narrowed to slender petiole. Inflorescence: head-like, axis compressed; bracts 3--9 mm, oblanceolate or spoon-shaped, spreading; pedicels 2--5 mm, 0.1--0.2 mm diam, spreading, straight, tip not curved. Flower: not inverted; hypanthium +- 1 mm; sepals 2--6 mm, oblanceolate, erect; corolla not conspicuously 2-lipped, divided 1/2 to base, white, tube bell-shaped, lobes spreading, elliptic or oblong, acute, 1.5--2 mm; filaments declined, curled up, 1--3 mm, few hairs at tip, appendage attached at base of filament, paddle-shaped, cells tube-like, blunt, 0.1 mm, anthers 0.5--0.6 mm; ovary 1/2--2/3-inferior. Fruit: 4--5 mm, base acute, tip dome-like; circumscissile. Seed: 0.5--0.7 mm, widely elliptic, pitted.
Ecology: Sandy or gravelly soils; Elevation: 650--1900 m. Bioregional Distribution: se SCoRO, s SCoRI (Caliente Range), ne WTR (n Ventura Co.), DMoj (San Bernardino Co.). Flowering Time: Apr--May
Synonyms: Parishella californica A. Gray
eFlora Treatment Author: Nancy R. Morin
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Previous taxon: Nemacladus calcaratus
Next taxon: Nemacladus capillaris

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botanical illustration including Nemacladus californicus


Citation for this treatment: Nancy R. Morin 2017. Nemacladus californicus, in Jepson Flora Project (eds.) Jepson eFlora,, accessed on May 25, 2017.

Citation for the whole project: Jepson Flora Project (eds.) 2017. Jepson eFlora,, accessed on May 25, 2017.

Geographic subdivisions for Nemacladus californicus:
se SCoRO, s SCoRI (Caliente Range), ne WTR (n Ventura Co.), DMoj (San Bernardino Co.).
Markers link to CCH specimen records. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues. Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.