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Nemacladus calcaratus

Higher Taxonomy
Family: CampanulaceaeView DescriptionDichotomous Key

Habit: Annual to perennial herb [tree]. Leaf: generally cauline, generally simple, generally alternate, petioled or not; stipules 0. Inflorescence: cyme, panicle, raceme, spike, or flowers 1; terminal or in axils of leaf-like or reduced bracts. Flower: bisexual, cleistogamous or open, radial or bilateral, inverted (pedicel twisted 180°) or not; hypanthium generally present, +- fused to ovary; sepals generally 5; corolla radial to 2-lipped, petals generally fused, tube deeply divided on 1 side or not, lobes generally 5; stamens 5, free or +- fused (anthers, filaments fused into tube or filaments fused above middle); ovary inferior or 1/2 inferior (superior in fruit), chambers 1--3, placentas axile or parietal, ovules many, style generally 1, 2--5-branched. Fruit: generally capsule, open on sides or top by pores or short valves. Seed: many.
Genera In Family: +- 90 genera, +- 2500 species: worldwide. Note: Some cultivated for ornamental (Campanula, Jasione, Lobelia). Subfamilies sometimes treated as families. Positions of flower parts given after flowering inversion, if any. Parishella moved to Nemacladus.
eFlora Treatment Author: Nancy R. Morin, except as noted
Scientific Editor: Thomas J. Rosatti.
Genus: NemacladusView DescriptionDichotomous Key

Habit: Annual, from taproot. Stem: prostrate, decumbent, or erect; base generally +- brown or +- purple; branches 0 or below middle. Leaf: basal; petiole short or 0. Inflorescence: +- raceme-like; bract 1 per flower, small; pedicel generally thread-like. Flower: inverted or not; sepals linear to triangular; corolla +- radial or 2-lipped, lobes 5; filaments free at base, fused into tube around style distally, appendages attached to a stalk or directly on 2 adjacent filaments, each with 2--12 cells, anthers free, all alike; ovary superior to 1/2 inferior, hemispheric to obconic, nectary glands 3, mounded or donut-like, on free part of ovary, stigma 2-lobed, papillate. Fruit: generally > hypanthium, hemispheric to fusiform, top pointed or rounded, chambers 2; open at top generally by 2 valves (or circumscissile). Seed: elliptic to oblong.
Species In Genus: 18 species: southwestern United States, northwestern Mexico. Etymology: (Greek: thread-like branch) Note: In descriptions, "filaments" including both free and fused parts thereof.
Unabridged Note: Taxonomic changes from TJM (1993) based on ITS, atpB, morphology. Parishella californica nested within Nemacladus, in which it is here treated.

Nemacladus calcaratus Morin
Habit: Erect, 2--4 cm, branches from base or 0.5--1 cm above. Stem: ascending; base dull, red-brown. Leaf: 2--5 mm, narrowly lanceolate to +- spoon-shaped or elliptic, remotely toothed, hairy, narrowed to wide petiole. Inflorescence: axis +- zigzag; bracts 1--1.2(2) mm, lance-ovate, clasping pedicel at base, reflexed above; pedicels 10--12 mm, +- 0.1 mm diam, spreading, straight or arched. Flower: not inverted; hypanthium +- 0.5 mm; sepals 0.8--1.5 mm, lanceolate, lowest reflexed, others ascending to erect; corolla divided 2/3 to base, white with pink or red midvein, tube widely cup-shaped, lobes all held on adaxial side of flower, bases of 2 lowest fused into spur; filaments slightly upcurved, 1.8--2 mm, tip +- curved, long-hairy, appendages stalk-like, attached at base of ovary, cells narrow, pointed, anthers 0.3--0.4 mm; ovary +- 1/2-inferior. Fruit: +- 3 mm, base, tip rounded. Seed: 1--1.2 mm, oblong, impressed vertical lines crossed by fine transverse lines.
Ecology: Decomposed granite flats; Elevation: 1900--2100 m. Bioregional Distribution: s SNH (Chimney Creek). Flowering Time: Jun
eFlora Treatment Author: Nancy R. Morin
Jepson Online Interchange
Listed on CNPS Rare Plant Inventory

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Next taxon: Nemacladus californicus

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Citation for this treatment: Nancy R. Morin 2016. Nemacladus calcaratus, in Jepson Flora Project (eds.) Jepson eFlora,, accessed on October 26, 2016.

Citation for the whole project: Jepson Flora Project (eds.) 2016. Jepson eFlora,, accessed on October 26, 2016.

Geographic subdivisions for Nemacladus calcaratus:
s SNH (Chimney Creek).
Markers link to CCH specimen records. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues. Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.