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Johanneshowellia puberula

Higher Taxonomy
Family: PolygonaceaeView DescriptionDichotomous Key

Habit: Annual to shrub [tree]. Stem: nodes swollen or not. Leaf: simple, basal or cauline, generally alternate; ocreae present or 0, generally scarious, persistent or not. Inflorescence: flower clusters in axillary to terminal cyme-, panicle-, raceme-, spike-, umbel- or head-like arrangements, entire inflorescence or main inflorescence branches generally subtended by bracts ("inflorescence bracts"); peduncles present or 0; flower clusters in Eriogoneae-Eriogonoideae subtended by involucre of >= 1 free or +- fused, sometimes awn-tipped bracts ("involucre bracts") or, in Polygonoideae and rarely in Eriogonoideae, not (if bracts completely fused, involucre "tubular"); pedicels in Eriogoneae each often subtended by 2 free, transparent, linear bractlets or in Polygonoideae all subtended by 2+ fused, membranous, wide bractlets. Flower: generally bisexual, small, 1--200 per node; perianth parts 2--6, generally in 2 whorls, free or basally fused, generally petal-like, often +- concave adaxially, often darker at midvein, often turning +- red or +- brown in age; stamens [1]3 or 6--9 in 2 whorls; ovary superior, 1-chambered, ovule 1, styles 1--3. Fruit: achenes, included in or exserted from perianth, generally 3-angled, ovoid or elliptic, generally glabrous.
Genera In Family: 48 genera, +- 1200 species: worldwide, especially northern temperate; some cultivated for food (Coccoloba, sea-grape; Fagopyrum, Rheum, Rumex) or ornamental (Antigonon, lovechain; Coccoloba; Muehlenbeckia; Persicaria; Polygonum), a few timbered (Coccoloba; Triplaris). Several (Emex; Fallopia; Persicaria; Polygonum; Rumex) are weeds. Note: Treatment of genera in Eriogonoideae based on monographic work of James L. Reveal. Involucre number throughout is number (1--many) per ultimate grouping, at tips of ultimate branches; flower number is per flower cluster or involucre, unless otherwise stated. Fagopyrum esculentum Moench not naturalized, considered an historical waif (or garden weed +- presently), therefore not treated.
eFlora Treatment Author: Mihai Costea, except as noted
Scientific Editor: Thomas J. Rosatti, Bruce G. Baldwin.
Genus: JohanneshowelliaView Description 

Habit: Annual, weakly erect to spreading, silky-puberulent. Leaf: basal; blades obovate to +- round [+- reniform], awns 0; ocreae 0. Inflorescence: terminal, cyme-like; peduncles 0; involucre bracts obscure, 4--7 in a spiral, narrow, awns 0. Flower: 4--7, glabrous, smooth or papillate; perianth white or pale +- yellow to rose or red, lobes 6, entire; stamens 9. Fruit: light brown, obconic, glabrous; embryo curved.
Species In Genus: 2 species: western North America. Etymology: (John Thomas Howell, 1903--1994, Eriogonum scholar, assistant to Alice Eastwood and her successor as Curator of Botany of the California Academy of Sciences)
eFlora Treatment Author: James L. Reveal & Thomas J. Rosatti
Reference: Reveal 2004 Brittonia 56:299--306

Johanneshowellia puberula (S. Watson) Reveal
Habit: Plant erect, 0.5--3 dm. Leaf: blades 0.5--1.5 cm. Inflorescence: involucre bracts 0.5--1.2 mm, 0.1--0.3 mm wide, outermost 2--3-lobed. Flower: 1--2 mm, minutely papillate; perianth white or pale yellow to rose or red. Fruit: 1--1.5 mm.
Ecology: Sand; Elevation: (500)800--2800 m. Bioregional Distribution: n DMtns (Cottonwood Mtns, Death Valley region, Inyo Co.); Distribution Outside California: to southwestern Utah. Flowering Time: May--Sep
Synonyms: Eriogonum puberulum S. Watson; Eriogonum puberulum var. venosum S. Stokes;
eFlora Treatment Author: James L. Reveal & Thomas J. Rosatti
Reference: Reveal 2004 Brittonia 56:299--306
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botanical illustration including Johanneshowellia puberula


Citation for this treatment: James L. Reveal & Thomas J. Rosatti 2012, Johanneshowellia puberula, in Jepson Flora Project (eds.) Jepson eFlora,, accessed on January 17, 2018.

Citation for the whole project: Jepson Flora Project (eds.) 2018, Jepson eFlora,, accessed on January 17, 2018.

Johanneshowellia puberula
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© 2011 Dana York

More photos of Johanneshowellia puberula in CalPhotos

Geographic subdivisions for Johanneshowellia puberula:
n DMtns (Cottonwood Mtns, Death Valley region, Inyo Co.);
Markers link to CCH specimen records. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues. Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.