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Horkelia californica var. californica

Higher Taxonomy
Family: RosaceaeView DescriptionDichotomous Key

Common Name: ROSE FAMILY
Habit: Annual to tree, glandular or not. Leaf: simple to palmately or pinnately compound, generally alternate; stipules free to fused (0), persistent to deciduous. Inflorescence: cyme, raceme, panicle, cluster, or flowers 1; bractlets on pedicel ("pedicel bractlets") generally 0--3(many), subtended by bract or generally not. Flower: generally bisexual, radial; hypanthium free or fused to ovary, saucer- to funnel-shaped, subtending bractlets ("hypanthium bractlets") 0--5, alternate sepals; sepals generally 5; petals generally 5, free; stamens (0,1)5--many, anther pollen sacs generally 2; pistils (0)1--many, simple or compound, ovary superior to inferior, styles 1--5. Fruit: 1--many per flower, achene (fleshy-coated or not), follicle, drupe, or pome with generally papery core, occasionally drupe-like with 1--5 stones. Seed: generally 1--5 (per fruit, not per flower).
Genera In Family: 110 genera, +- 3000 species: worldwide, especially temperate; many cultivated for ornamental, fruit, especially Cotoneaster, Fragaria, Malus, Prunus, Pyracantha, Rosa, Rubus. Note: Number of teeth is per leaf or leaflet, not per side of leaf or leaflet, except in Drymocallis.
eFlora Treatment Author: Daniel Potter & Barbara Ertter, family description, key to genera, treatment of genera by Daniel Potter, except as noted
Scientific Editor: Daniel Potter, Thomas J. Rosatti.
Genus: HorkeliaView DescriptionDichotomous Key

Habit: Perennial herb, generally +- glandular, generally resinous-smelling; caudex generally branched. Stem: generally ascending to erect. Leaf: generally basal, odd-1-pinnately compound, generally +- flat; cauline alternate, reduced upward; leaflets 2--15 per side, uppermost lateral generally +- fused with terminal. Inflorescence: cyme; pedicels generally straight, bractlets 0. Flower: hypanthium cup-like, +- flat-bottomed, width +- 2 × length, bractlets 5, generally 2/3 sepals; sepals often reflexed; petals generally +- = sepal, blunt, white; stamens 10, filaments +- flat, often forming a tube; pistils 2--many, ovary superior, style attached below fruit tip, +- thicker at base. Fruit: achene.
Species In Genus: 20 species: western North America. Etymology: (J. Horkel, German plant physiologist, 1769--1846) Note: Many attractive to bees; data apply to basal leaves, pressed hypanthia.
eFlora Treatment Author: Barbara Ertter
Reference: Ertter & Reveal 2007 Novon 17:315--325
Species: Horkelia californicaView Description 

Habit: Plant clumped, green. Stem: 10--120 cm. Leaf: stipules entire; leaflets +- separated, +- ovate to round, +- hairy. Inflorescence: open, of few to many separate flowers and few-flowered clusters; pedicels generally 1--20 mm. Flower: hypanthium width 4--10 mm, 1--2 × length, bractlets +- 4--6 mm, generally +- 2 mm wide, ovate, often toothed; sepals generally 4--6 mm; petals 3--8 mm, generally oblanceolate to elliptic; anthers 0.8--1.8 mm; pistils generally > 50. Fruit: +- 1 mm.
Note: Often confused with Drymocallis species (terminal leaflets distinct). Varieties intergrade.

Horkelia californica Cham. & Schltdl. var. californica
Leaf: 8--40 cm; sheathing base +- hairy; leaflets 4--9 per side, 10--40 mm, toothed, generally few-lobed +- 1/2 to base; terminal leaflet 10--40 mm. Flower: hypanthium inner wall +- hairy, bractlets generally toothed; sepals red-mottled inside; filaments generally 1.5--3 mm, base 0.5--1 mm wide; style generally 3--4 mm. Chromosomes: n=28.
Ecology: Grassy openings, edges of coastal scrub, especially northern slopes; Elevation: < 400 m. Bioregional Distribution: NCo, CCo. Flowering Time: Mar--Sep
Synonyms: Horkelia californica subsp. californica;
eFlora Treatment Author: Barbara Ertter
Reference: Ertter & Reveal 2007 Novon 17:315--325
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botanical illustration including Horkelia californica var. californica


Citation for this treatment: Barbara Ertter 2012, Horkelia californica var. californica, in Jepson Flora Project (eds.) Jepson eFlora,, accessed on October 23, 2018.

Citation for the whole project: Jepson Flora Project (eds.) 2018, Jepson eFlora,, accessed on October 23, 2018.

Horkelia californica var. californica
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© 2016 Neal Kramer
Horkelia californica var. californica
click for enlargement
© 2016 Neal Kramer
Horkelia californica var. californica
click for enlargement
© 2016 Neal Kramer
Horkelia californica var. californica
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© 2014 Vernon Smith

More photos of Horkelia californica var. californica in CalPhotos

Geographic subdivisions for Horkelia californica var. californica:
NCo, CCo.
Markers link to CCH specimen records. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues. Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa, if there are more than 1 infraspecific taxon in CA.
Blue line denotes eFlora flowering time.