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Ericameria nauseosa var. bernardina
BERNARDINA RABBITBRUSH

Higher Taxonomy
Family: Asteraceae (Compositae)View DescriptionDichotomous Key

Common Name: SUNFLOWER FAMILY
Habit: Annual to tree. Leaf: basal and/or cauline, alternate, opposite, rarely whorled, simple to 2+ × compound. Inflorescence: 1° inflorescence a head, resembling a flower, of several types (see below), 1--many in generally +- cyme-like cluster; each head generally with +- calyx-like involucre of 1--many series of phyllaries (involucre bracts); receptacle of head flat to conic or columnar, paleate (bearing paleae = receptacle bracts) or epaleate; flowers 1--many per head. Flower: bisexual, unisexual, or sterile, +- small, of several types (see below); calyx 0 or modified into +- persistent pappus of bristles, scales, and/or awns; corolla radial or bilateral (0), lobes generally (0)3--5; stamens 4--5, filaments generally free, generally fused to corolla at tube/throat junction, anthers generally fused into cylinder around style, anther base generally rounded or cordate (deeply sagittate or with tail-like appendages), tip (= flattened appendage) generally projecting beyond pollen sac; pistil 1, 2-carpeled, ovary inferior, 1-chambered, 1-seeded, placenta basal, style 1, tip generally +- 2-branched (except in some staminate disk flowers), branch tips truncate or generally bearing +- brush-like appendages; stigmas 2, generally on adaxial faces of style branches. Fruit: achene (also called a cypsela) (drupe in Chrysanthemoides), cylindric to ovoid, sometimes compressed, generally deciduous with pappus attached.
Genera In Family: +- 1500 genera, 23000 species: worldwide, many habitats. Note: Flower and head types differ in form and sexual condition. A disk flower has a generally radial corolla, with a cylindric tube, expanded throat, and generally 5 lobes. Disk flowers are generally bisexual and fertile but occasionally staminate with reduced ovaries. Discoid heads comprise only disk flowers. A radiant head is a variant of a discoid head, with peripheral disk flower corollas expanded, often bilateral. A ray flower corolla is bilateral, generally with a slender tube and flattened petal-like ray (single lip composed of generally 3 lobes). Ray flowers are generally pistillate or sterile (occasionally lacking styles). Radiate heads have peripheral ray flowers and central disk flowers. Disciform heads superficially resemble discoid heads, with pistillate or sterile flowers that lack rays, together with or separate from disk flowers. A ligulate flower is bisexual, with a bilateral, generally ephemeral corolla and 5-lobed ligule. Liguliflorous heads comprise only ligulate flowers. See glossary p. 31 for illustrations of family characteristics. Echinops sphaerocephalus L., Gaillardia aristata Pursh, Gaillardia pulchella Foug., Hymenothrix loomisii S.F. Blake, Tagetes erecta L., Thelesperma megapotamicum (Spreng.) Kuntze are waifs. Melampodium perfoliatum Kunth, historic urban waif. Ageratum conyzoides L., Guizotia abyssinica (L. f.) Cass., Santolina chamaecyparisus L., orth. var. are rare or uncommon escapes from cultivation. Dyssodia papposa, Ismelia carinata (Schousb.) Sch. Bip. [Chrysanthemum carinatum Schousb.], Mantisalca salmantica (L.) Briq. & Cavill. are historical or extirpated waifs in California. Inula helenium L. not documented in California. Taxa of Aster in TJM (1993) treated here in Almutaster, Eucephalus, Eurybia, Ionactis, Oreostemma, Sericocarpus, Symphyotrichum; Chamomilla in Matricaria; Cnicus in Centaurea; Conyza in Erigeron and Laennecia; Dugaldia in Hymenoxys; Erechtites in Senecio; Hymenoclea in Ambrosia; Lembertia in Monolopia; Osteospermum ecklonis in Dimorphotheca; Picris echioides in Helminthotheca; Prionopsis in Grindelia; Raillardiopsis in Anisocarpus and Carlquistia; Schkuhria multiflora in Bahia; Trimorpha in Erigeron; Venidium in Arctotis; Whitneya in Arnica. Taxa of Arida in TJM2 (2012) treated here in Leucosyris.
Unabridged Note: Largest family of vascular plants in California and of eudicots globally.
eFlora Treatment Author: David J. Keil, except as noted
Scientific Editor: David J. Keil, Bruce G. Baldwin.
Genus: EricameriaView DescriptionDichotomous Key

Common Name: GOLDENBUSH, RABBITBRUSH
Habit: Subshrub, shrub, <= 5 m, resinous, generally gland-dotted. Stem: generally ascending to erect, glabrous to woolly; often resin-dotted and resin-coated, stalked-glandular in some. Leaf: alternate, thread-like to elliptic or wedge-shaped, entire. Inflorescence: heads radiate or discoid, 1 or variously clustered; involucre cylindric, obconic, or bell-shaped; phyllaries in 2--7 series, +- lanceolate to ovate, generally resinous, persistent when dry, tips erect to recurved, obtuse to acuminate or tailed, midrib often thickened with a resin gland; +- flat to convex, pitted, epaleate. Ray Flower: 0--30; corolla 2--12 mm, generally yellow (white). Disk Flower: 4--70+; corolla 3--11 mm, generally yellow (white); anther tips narrowly triangular to awl-like; style tips lanceolate to awl-shaped. Fruit: 2--8 mm, cylindric to ellipsoid or +- obconic, generally angled, ribbed; pappus of minutely barbed bristles in 1 series.
Species In Genus: +- 36 species: western North America. Etymology: (Greek: golden shrub) Note: Most species flower summer/fall. Some hybridization among species.
eFlora Treatment Author: Lowell E. Urbatsch
Species: Ericameria nauseosaView Description 

Common Name: RUBBER RABBITBRUSH
Habit: Plant 2--28 dm; strongly scented. Stem: spreading or ascending to erect, +- flexible, very leafy or leafless at flower, +- white to green, loosely to densely tomentose. Leaf: 10--70 mm, thread-like to narrowly (ob)lanceolate, glabrous to tomentose, gland-dotted or not. Inflorescence: heads discoid, many in dense, flat-topped or rounded to +- panicle-like clusters; peduncle 1--20 mm, bracts generally 0(1--5, scale-like); involucre 6--14 mm, 2--4 mm wide, cylindric; phyllaries 10--31, graduated in 3--5 series, generally in vertical ranks (appearing spiraled), +- lanceolate to ovate, generally +- strongly keeled, firm, dry, paper-like, obtuse to acute, tips long-tapered, erect to recurved. Ray Flower: 0. Disk Flower: generally 5; corolla 6--13 mm, lobes generally glabrous (long-soft-hairy); style appendage < to > stigma. Fruit: 3--10 mm, generally hairy; pappus 3--13 mm, +- white. Chromosomes: 2n=18.
Note: Highly variable; 13 more varieties in western North America, northwestern Mexico.

Ericameria nauseosa var. bernardina (H.M. Hall) G.L. Nesom & G.I. Baird
NATIVE
Habit: Plant 5--15 dm, white-tomentose to green. Stem: leafy at flower, closely tomentose, +- yellow-green, becoming +- white. Leaf: 25--60 mm, +- narrowly lanceolate. Inflorescence: involucre 10--14 mm, glabrous to hairy. Flower: corolla 10--12 mm, lobes 1.7--2.3 mm, ascending to spreading; style appendage > stigma.
Ecology: Open yellow-pine forest; Elevation: 1200--3000 m. Bioregional Distribution: s SN, TR, n PR (incl SnJt), W&I; Distribution Outside California: northern Baja California. Flowering Time: Aug--Oct Note: Much like robust Ericameria nauseosa var. speciosa. Sometimes also confused with Ericameria nauseosa var. oreophila, Ericameria nauseosa var. mohavensis.
Synonyms: Chrysothamnus nauseosus subsp. bernardinus (H.M. Hall) H.M. Hall & Clem.
eFlora Treatment Author: Lowell E. Urbatsch
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Citation for this treatment: Lowell E. Urbatsch 2016. Ericameria nauseosa var. bernardina, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=80500, accessed on August 23, 2016.

Citation for the whole project: Jepson Flora Project (eds.) 2016. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on August 23, 2016.


Ericameria nauseosa var. bernardina
click for enlargement
© 2015 Keir Morse
Ericameria nauseosa var. bernardina
click for enlargement
© 2015 Keir Morse
Ericameria nauseosa var. bernardina
click for enlargement
© 2013 Steve Matson
Ericameria nauseosa var. bernardina
click for enlargement
© 2013 Steve Matson
Ericameria nauseosa var. bernardina
click for enlargement
© 2015 Keir Morse
Ericameria nauseosa var. bernardina
click for enlargement
© 2013 Steve Matson

More photos of Ericameria nauseosa var. bernardina in CalPhotos


Geographic subdivisions for Ericameria nauseosa var. bernardina:
s SN, TR, n PR (incl SnJt), W&I;
Markers link to CCH specimen records. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues. Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.