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Vascular Plants of California
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Harmonia doris-nilesiae

Higher Taxonomy
Family: Asteraceae (Compositae)View DescriptionDichotomous Key
Habit: Annual to tree. Leaf: basal and/or cauline, alternate, opposite, rarely whorled, simple to 2+ × compound. Inflorescence: 1° inflorescence a head, resembling a flower, of several types (see below), 1--many in generally +- cyme-like cluster; each head generally with +- calyx-like involucre of 1--many series of phyllaries (involucre bracts); receptacle of head flat to conic or columnar, paleate (bearing paleae = receptacle bracts) or epaleate; flowers 1--many per head. Flower: bisexual, unisexual, or sterile, +- small, of several types (see below); calyx 0 or modified into +- persistent pappus of bristles, scales, and/or awns; corolla radial or bilateral (0), lobes generally (0)3--5; stamens 4--5, filaments generally free, generally fused to corolla at tube/throat junction, anthers generally fused into cylinder around style, anther base generally rounded or cordate (deeply sagittate or with tail-like appendages), tip (= flattened appendage) generally projecting beyond pollen sac; pistil 1, 2-carpeled, ovary inferior, 1-chambered, 1-seeded, placenta basal, style 1, tip generally +- 2-branched (except in some staminate disk flowers), branch tips truncate or generally bearing +- brush-like appendages; stigmas 2, generally on adaxial faces of style branches. Fruit: achene (also called a cypsela) (drupe in Chrysanthemoides), cylindric to ovoid, sometimes compressed, generally deciduous with pappus attached.
Genera In Family: +- 1500 genera, 23000 species: worldwide, many habitats. Note: Flower and head types differ in form and sexual condition. A disk flower has a generally radial corolla, with a cylindric tube, expanded throat, and generally 5 lobes. Disk flowers are generally bisexual and fertile but occasionally staminate with reduced ovaries. Discoid heads comprise only disk flowers. A radiant head is a variant of a discoid head, with peripheral disk flower corollas expanded, often bilateral. A ray flower corolla is bilateral, generally with a slender tube and flattened petal-like ray (single lip composed of generally 3 lobes). Ray flowers are generally pistillate or sterile (occasionally lacking styles). Radiate heads have peripheral ray flowers and central disk flowers. Disciform heads superficially resemble discoid heads, with pistillate or sterile flowers that lack rays, together with or separate from disk flowers. A ligulate flower is bisexual, with a bilateral, generally ephemeral corolla and 5-lobed ligule. Liguliflorous heads comprise only ligulate flowers. See glossary p. 31 for illustrations of family characteristics. Echinops sphaerocephalus L., Gaillardia aristata Pursh, Gaillardia pulchella Foug., Hymenothrix loomisii S.F. Blake, Tagetes erecta L., Thelesperma megapotamicum (Spreng.) Kuntze are waifs. Melampodium perfoliatum Kunth, historic urban waif. Ageratum conyzoides L., Guizotia abyssinica (L. f.) Cass., Santolina chamaecyparisus L., orth. var. are rare or uncommon escapes from cultivation. Dyssodia papposa, Ismelia carinata (Schousb.) Sch. Bip. [Chrysanthemum carinatum Schousb.], Mantisalca salmantica (L.) Briq. & Cavill. are historical or extirpated waifs in California. Inula helenium L. not documented in California. Taxa of Aster in TJM (1993) treated here in Almutaster, Eucephalus, Eurybia, Ionactis, Oreostemma, Sericocarpus, Symphyotrichum; Chamomilla in Matricaria; Bahia in Hymenothrix; Cnicus in Centaurea; Conyza in Erigeron and Laennecia; Dugaldia in Hymenoxys; Erechtites in Senecio; Hymenoclea in Ambrosia; Lembertia in Monolopia; Osteospermum ecklonis in Dimorphotheca; Picris echioides in Helminthotheca; Prionopsis in Grindelia; Raillardiopsis in Anisocarpus and Carlquistia; Schkuhria multiflora in Picradeniopsis; Trimorpha in Erigeron; Venidium in Arctotis; Whitneya in Arnica. Amauriopsis in TJM2 (2012) treated here in Hymenothrix; Arida in Leucosyris; Bahia in Picradeniopsis.
Unabridged Note: Largest family of vascular plants in California and of eudicots globally.
eFlora Treatment Author: David J. Keil, except as noted
Scientific Editor: David J. Keil, Bruce G. Baldwin.
Genus: HarmoniaView DescriptionDichotomous Key

Habit: Annual 0.5--4 dm. Stem: erect. Leaf: proximal opposite, distal alternate, sessile, linear, entire or toothed, generally coarsely hairy, sometimes also minutely stalked-glandular, glands generally black, sometimes +- yellow. Inflorescence: heads radiate, 1 or in loose, +- umbel-like to flat-topped clusters; involucre obovoid to obconic, 2--5+ mm diam; phyllaries 3--8 in 1 series, lanceolate to oblanceolate, each enclosing a ray ovary, falling with fruit; receptacle flat to convex, glabrous or minutely bristly, paleae in 1 series between ray and disk flowers, free or fused, phyllary-like (more scarious), deciduous. Ray Flower: generally 3--8; corolla bright yellow, ray fan-shaped to obovate. Disk Flower: 7--30, bisexual or staminate, sometimes in same head; corolla bright yellow, tube < throat, lobes deltate; anthers +- yellow to +- brown, tips ovate-dentate to hemispheric; styles glabrous proximal to branches, tips awl-shaped, densely bristly.; anther bases sagittate. Fruit: ray fruit black, round in ×-section to +- compressed, weakly arched, bowed abaxially or not, beaked or beakless, glabrous, pappus 0 or of 3--12, lanceolate to awl-shaped, fringed to plumose scales; disk fruit black, +- club-shaped, +- round, glabrous or hairy, pappus of 7--11, generally linear to lanceolate or awl-shaped, fringed or plumose scales.
Species In Genus: 5 species: northwestern California. Etymology: (Harvey Monroe Hall, California botanist, 1874--1932)
eFlora Treatment Author: Bruce G. Baldwin
Reference: Baldwin & Strother 2006 FNANM 21:297--299
Unabridged Reference: Baldwin 2001 Madroño 48:293--297
Harmonia doris-nilesiae (T.W. Nelson & J.P. Nelson) B.G. Baldwin
Habit: Plant 9--40 cm. Stem: proximal, unbranched part of central stem < or > branches supporting heads. Leaf: +- evenly distributed on stems. Inflorescence: heads generally erect in bud and fruit; phyllaries 4--8, densely white-soft-hairy near folded edges. Ray Flower: 4--8; ray 2.5--7 mm. Disk Flower: 8--20, staminate. Fruit: ray fruit strongly bowed abaxially, beak 0.4--0.5 mm, pappus to 0.9 mm; disk fruit 0, pappus of 7--10, lanceolate-long-tapered to awl-shaped, plumose scales 0.2--0.9 mm. Chromosomes: 2n=18.
Ecology: Serpentine slopes; Elevation: 800--1600 m. Bioregional Distribution: s KR. Flowering Time: May--Jun
Synonyms: Madia doris-nilesiae T.W. Nelson & J.P. Nelson
Jepson eFlora Author: Bruce G. Baldwin
Reference: Baldwin & Strother 2006 FNANM 21:297--299
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Botanical illustration including Harmonia doris-nilesiae

botanical illustration including Harmonia doris-nilesiae


Citation for this treatment: Bruce G. Baldwin 2012, Harmonia doris-nilesiae, in Jepson Flora Project (eds.) Jepson eFlora,, accessed on October 18, 2019.

Citation for the whole project: Jepson Flora Project (eds.) 2019, Jepson eFlora,, accessed on October 18, 2019.

Harmonia doris-nilesiae
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© 2017 Julie Kierstead Nelson
Harmonia doris-nilesiae
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© 2013 Julie Kierstead Nelson
Harmonia doris-nilesiae
click for enlargement
© 2017 Julie Kierstead Nelson
Harmonia doris-nilesiae
click for enlargement
© 2019 Steve Matson
Harmonia doris-nilesiae
click for enlargement
© 2019 Steve Matson
Harmonia doris-nilesiae
click for enlargement
© 2019 Steve Matson

More photos of Harmonia doris-nilesiae in CalPhotos

Geographic subdivisions for Harmonia doris-nilesiae:
s KR.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).


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Data provided by the participants of the  Consortium of California Herbaria.
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Blue markers indicate specimens that map to one of the expected Jepson geographic subdivisions (see left map). Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa, if there are more than 1 infraspecific taxon in CA.
Blue line denotes eFlora flowering time.