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Bolboschoenus robustus
SEACOAST BULRUSH

Higher Taxonomy
Family: CyperaceaeView DescriptionDichotomous Key

Common Name: SEDGE FAMILY
Habit: Annual, perennial herb, often rhizomed or stoloned, often of wet open places; roots fibrous; monoecious, dioecious, or flowers bisexual. Stem: generally 3-sided, generally solid. Leaf: generally 3-ranked; base sheathing, sheath generally closed, ligule generally 0; blade (0 or) linear, parallel-veined. Inflorescence: spikelets generally arranged in head-, spike-, raceme-, or panicle-like inflorescences; flower generally sessile in axil of flower bract, enclosed in a sac-like structure (perigynium) or generally not. Flower: unisexual or bisexual, small, generally wind-pollinated; perianth 0 or generally bristle like; stamens generally 3, anthers attached at base, 4 chambered; ovary superior, chamber 1, ovule 1, style 2--3-branched. Fruit: achene, 2--3 sided.
Genera In Family: +- 100 genera, 5000 species: especially temperate. Note: Difficult; taxa differ in technical characters of inflorescence, fruit. In Carex and Kobresia, what appear to be individual pistillate flowers in fact are highly reduced inflorescences (whether or not the same applies to staminate flowers is still under debate). In some other works (e.g., FNANM) these are called spikelets, and they are treated as being arranged in spikes. Here and in TJM (1993), what appear to be individual pistillate flowers are called pistillate flowers in Carex (and they are treated as being arranged in spikelets), but spikelets in Kobresia (and they are treated as being arranged into spikes). Though internally inconsistent, the approach here is consistent with traditional usage, and reflects a preference for character states that may be determined in the field. Molecular, morphological, and embryological evidence indicates that Eriophorum crinigerum is to be segregated to a new genus, as Calliscirpus criniger (A. Gray) C.N. Gilmour et al., along with a second, newly described species, Calliscirpus brachythrix C.N. Gilmour et al. (Gilmour et al. 2013); key to genera modified by Peter W. Ball to include Calliscirpus.
eFlora Treatment Author: S. Galen Smith, except as noted
Scientific Editor: S. Galen Smith, Thomas J. Rosatti, Bruce G. Baldwin.
Genus: BolboschoenusView DescriptionDichotomous Key

Common Name: TUBEROUS BULRUSHES
Habit: Perennial herb, erect, 50--200 cm, rhizomed, tubers durable. Stem: simple, sharply 3-angled, glabrous or angles scabrous, evident internal air cavities 0, not hollow. Leaf: basal and cauline, 3-ranked; sheath closed, long; ligule 0; blade generally present, long, thin, flat, V-shaped near base, keeled abaxially, margin, keel +- scabrous. Inflorescence: 1, terminal, panicle- (or head-) like, appearing with leaves; branches often scabrous; inflorescence bracts like leaf blades, main 1 > inflorescence; spikelets +- ovate, not +- flat, flower bracts spiraled, >= 25, each with 1 flower in axil, +- ovate, membranous to papery, puberulent (glabrous in age), brown to +- colorless, tip notched 0.5--1 mm, generally with curved awn often broken off. Flower: bisexual; perianth of 3--6 bristles, <= fruit, +- straight, stout, barbed; stamens 3, anthers >= 1.5 mm; style 1, thread-like, base not enlarged; stigmas 2--3. Fruit: generally obovate, smooth, brown, mucronate; wall cells small, solid or large, hollow (under dissecting microscope).
Species In Genus: 7--15 species: temperate, subtropics. Etymology: (Greek, bolbos, a bulb, and schoenos a rush, reed, in reference to the tubers) Note: Intermediates (putative hybrids) between species cause major problems in classification, identification.
Unabridged Note: Fruit wall anatomy (+- easily seen with a dissecting microscope in a hand-made section) including diagnostic characters that are correlated with fruit buoyancy and persistence of bristles on shed fruit. Putative interspecific hybridization causes major taxonomic confusion; putative hybrids generally occur with their parents, are intermediate between them in all characters, often bear apparently normal fruit that vary in shape within one spikelet, and form persistent clones.

Bolboschoenus robustus (Pursh) Soják
NATIVE
Habit: Plant 50--150 cm. Stem: 4--8 mm diam. Leaf: sheath tip generally papery, veiny; widest blade 4--12 mm wide. Inflorescence: proximal inflorescence bract 2--7 mm wide; spikelets (1)5--25, <= 1/2 on branches, 10--30 mm, 6--10 mm wide; flower bracts 6--9 mm, awn 2--3 mm, base +- 0.5 mm wide. Flower: perianth bristles not attached to (or some loosely attached), +- 1/2 fruit; anthers orange, stigmas 3(2). Fruit: 2.7--3.5 mm, 1.8--2.8 mm wide, +- flat-3-sided or 2-sided, floating on water; mucro 0.1--0.3 mm; wall cells large, hollow.
Ecology: Local in California. Brackish to saline coastal marshes; Elevation: +- 0 m. Bioregional Distribution: GV (Suisun Marsh), CCo, SnFrB, SCo; Distribution Outside California: Maine to Florida; Mexico, South America. Flowering Time: Summer
Synonyms: Scirpus robustus Pursh; Schoenoplectus robustus (Pursh) M.T. Strong
Unabridged Note: In North America Pacific Coastal literature Bolboschoenus (Scirpus) maritimus often has been erroneously called Scirpus robustus. For hybrids, see Bolboschoenus fluviatilis, Bolboschoenus maritimus.
eFlora Treatment Author: S. Galen Smith
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Citation for this treatment: S. Galen Smith 2016. Bolboschoenus robustus, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=77450, accessed on May 29, 2016.

Citation for the whole project: Jepson Flora Project (eds.) 2016. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on May 29, 2016.


Bolboschoenus robustus
click for enlargement
© 2004 Steve Matson
Bolboschoenus robustus
click for enlargement
© 2004 Steve Matson
Bolboschoenus robustus
click for enlargement
© 2004 Steve Matson
Bolboschoenus robustus
click for enlargement
© 2004 Steve Matson
Bolboschoenus robustus
click for enlargement
© 2004 Steve Matson
Bolboschoenus robustus
click for enlargement
© 2004 Steve Matson

More photos of Bolboschoenus robustus in CalPhotos


Geographic subdivisions for Bolboschoenus robustus:
GV (Suisun Marsh), CCo, SnFrB, SCo;
Markers link to CCH specimen records. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues. Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.