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Holosteum umbellatum subsp. umbellatum

Higher Taxonomy
Family: CaryophyllaceaeView DescriptionDichotomous Key

Common Name: PINK FAMILY
Habit: Annual to perennial herb; rarely dioecious (Silene), taprooted or rhizome generally slender. Leaf: simple, generally opposite (subwhorled), entire, pairs at nodes often +- connected at bases; stipules generally 0; petiole generally 0. Inflorescence: generally cyme, generally open; flowers 1--many; involucre generally 0 (present in Dianthus, Petrorhagia). Flower: generally bisexual, radial; hypanthium often present but obscure; sepals (4)5, +- free or fused into a tube, margins generally scarious, more so on inner 2 or not, tube generally not scarious, awns generally 0; petals (4)5 or 0, generally tapered to base (or with claw long, limb expanded), entire to 2--several-lobed, limb generally without scale-like appendages adaxially, generally without ear-like lobes at base; stamens generally 10, generally fertile, generally free, generally from ovary base; nectaries 0 or 5; ovary superior, generally 1-chambered, placentas basal or free-central, styles 2--5 with 0 branches or 1 with 2--3 branches. Fruit: capsule or utricle (rarely +- dehiscent), generally sessile.
Genera In Family: 83 or 89 genera, 3000 species: widespread, especially arctic, alpine, temperate northern hemisphere; some cultivated (Agrostemma, Arenaria, Cerastium, Dianthus, Gypsophila, Lychnis, Sagina, Saponaria, Silene, Vaccaria). Note: Apetalous Caryophyllaceae can also be keyed in Rabeler & Hartman 2005 FNANM 5:5--8.
eFlora Treatment Author: Ronald L. Hartman & Richard K. Rabeler, except as noted
Scientific Editor: Thomas J. Rosatti.
Genus: HolosteumView Description 

Habit: Annual, ascending to erect, taprooted. Leaf: basal blades oblanceolate to spoon-shaped, short-petioled; cauline blades narrowly oblanceolate to oblong-ovate, sessile; vein 1. Inflorescence: terminal, umbel-like; flowers 3--12; involucre bracts 5--15, +- 0.3--1.8 mm; pedicels 0.5--2.5 cm. Flower: sepals 5, 2.5--4.5 mm, lanceolate to ovate, glabrous; petals 5, 3.3--5 mm, +- irregularly toothed; stamens 3--5; styles 3, 0.5--1.5 mm. Fruit: capsule, ovoid or ovoid-cylindric, straight; teeth 6, recurved. Seed: many, red-brown.
Species In Genus: 3--4 species: temperate Eurasia, northern Africa. Etymology: (Greek: all bone, humorous reference to frailty of pl)

Holosteum umbellatum L. subsp. umbellatum
Habit: Plant 5--25 cm. Stem: 1--many, unbranched, glandular-hairy +- near middle. Leaf: cauline 1--4 pairs in lower 1/2; blades 2--30 mm, glabrous or margin glandular-hairy. Inflorescence: bracts whorled, 1--2 mm, partly scarious; pedicels slender, reflexed in flower, erect in fruit. Flower: petals narrowly elliptic, white. Seed: 0.7--0.8 mm, compressed, with marginal ridge opposite elongate depression; tubercles low, rounded. Chromosomes: 2n=20,40.
Ecology: Uncommon. Disturbed areas, roadsides; Elevation: < 1270 m. Bioregional Distribution: KR, CaR, CCo?, MP; Distribution Outside California: to British Columbia, eastern North America; native to Europe; central and southwestern Asia; northern Africa. Flowering Time: Late winter--early spring
Unabridged Note: 4 other subspecies, native to Europe, central and southwestern Asia, northern Africa, introduced in South America, western Europe, southern Africa.
eFlora Treatment Author: Ronald L. Hartman & Richard K. Rabeler
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botanical illustration including Holosteum umbellatum subsp. umbellatum


Citation for this treatment: Ronald L. Hartman & Richard K. Rabeler 2016. Holosteum umbellatum subsp. umbellatum, in Jepson Flora Project (eds.) Jepson eFlora,, accessed on April 30, 2016.

Citation for the whole project: Jepson Flora Project (eds.) 2016. Jepson eFlora,, accessed on April 30, 2016.

Geographic subdivisions for Holosteum umbellatum subsp. umbellatum:
KR, CaR, CCo?, MP;
Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.