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|Family: Pteridaceae||View Description||Dichotomous Key|
|Genus: Pentagramma||View Description||Dichotomous Key|
|Species:||View Description||Dichotomous Key|
Habit: Rhizome tip, scales without exudate. Leaf: blade adaxially glabrous, not sticky, margins not recurved; abaxially with generally yellow to cream (white) exudate; proximal pinnae 2--6(11) cm; distal pinnae, pinnules on basal side of proximal-most pinnae pinnately lobed. Chromosomes: 2n=60, 90, 120.
Ecology: Common. Generally shaded, sometimes rocky or wooded areas; Elevation: < 2300 m. Bioregional Distribution: CA-FP, MP (caves in Lava Beds National Monument), SNE, DMtns; Distribution Outside California: to British Columbia, Idaho, Nevada, Utah, Baja California. Note: Plants with dense white (instead of yellow) exudate on abaxial blade surfaces have been treated by some as Pentagramma triangularis subsp. semipallida (J.T. Howell) Yatsk. et al.; however, such plants appear to be evolutionarily embedded within yellow exudate forms (Schuettpelz et al. 2015 Syst Bot 40:629--644). Chromosome counts from plants of both white-exudate ("semipallida") and yellow-exudate morphologies (Alt & Grant 1960 Brittonia 12:153--170; Windham & Yatskievych 2003 Amer J Bot 90:1788--1800). Based on spore size, polyploids common, including likely triploids, tetraploids, and hexaploids (Alt & Grant 1960 Brittonia 12:153--170; Schuettpelz et al. 2015 Syst Bot 40:629--644; Smith et al. 1971 Amer J Bot 58:292--299).
Synonyms: Pentagramma triangularis subsp. semipallida (J.T. Howell) Yatsk., Windham & E. Wollenw.; Pityrogramma triangularis (Kaulf.) Maxon var. triangularis
Unabridged Note: Until the study of Schuettpelz et al. (2015 Syst Bot 40:629--644) Pentagramma triangularis generally treated to include P. glanduloviscida, P. maxonii, P. rebmanii, and P. viscosa. As treated here, P. triangularis restricted to plants with glabrous adaxial leaf surfaces. These plants form an evolutionary cluster, separate from other lineages of Pentagramma, regardless of whether their abaxial leaf surfaces have yellow exudate (often recognized as P. triangularis subsp. triangularis) or white exudate (often recognized as P. triangularis subsp. semipallida). These two exudate forms---yellow versus white---not well differentiated from each other evolutionarily (Schuettpelz et al. 2015 Syst Bot 40:629--644). Even as treated here, P. triangularis is the most wide-ranging species in the genus, with extensive morphological, chromosomal and chemical variation (Alt & Grant 1960 Brittonia 12:153--170).
eFlora Treatment Author: Ruth E.B. Kirkpatrick, Alan R. Smith & Thomas Lemieux
Jepson Online Interchange
Previous taxon: Pentagramma rebmanii
Next taxon: Pentagramma viscosa
Citation for this treatment: Ruth E.B. Kirkpatrick, Alan R. Smith & Thomas Lemieux 2017. Pentagramma triangularis, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=77119, accessed on February 19, 2017.
Citation for the whole project: Jepson Flora Project (eds.) 2017. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on February 19, 2017.