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    Vascular Plants of California
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Adenophyllum cooperi

Higher Taxonomy
Family: Asteraceae (Compositae)View DescriptionDichotomous Key

Habit: Annual to tree. Leaf: basal and/or cauline, alternate, opposite, rarely whorled, simple to 2+ × compound. Inflorescence: 1° inflorescence a head, resembling a flower, of several types (see below), 1--many in generally +- cyme-like cluster; each head generally with +- calyx-like involucre of 1--many series of phyllaries (involucre bracts); receptacle of head flat to conic or columnar, paleate (bearing paleae = receptacle bracts) or epaleate; flowers 1--many per head. Flower: bisexual, unisexual, or sterile, +- small, of several types (see below); calyx 0 or modified into +- persistent pappus of bristles, scales, and/or awns; corolla radial or bilateral (0), lobes generally (0)3--5; stamens 4--5, filaments generally free, generally fused to corolla at tube/throat junction, anthers generally fused into cylinder around style, anther base generally rounded or cordate (deeply sagittate or with tail-like appendages), tip (= flattened appendage) generally projecting beyond pollen sac; pistil 1, 2-carpeled, ovary inferior, 1-chambered, 1-seeded, placenta basal, style 1, tip generally +- 2-branched (except in some staminate disk flowers), branch tips truncate or generally bearing +- brush-like appendages; stigmas 2, generally on adaxial faces of style branches. Fruit: achene (also called a cypsela) (drupe in Chrysanthemoides), cylindric to ovoid, sometimes compressed, generally deciduous with pappus attached.
Genera In Family: +- 1500 genera, 23000 species: worldwide, many habitats. Note: Flower and head types differ in form and sexual condition. A disk flower has a generally radial corolla, with a cylindric tube, expanded throat, and generally 5 lobes. Disk flowers are generally bisexual and fertile but occasionally staminate with reduced ovaries. Discoid heads comprise only disk flowers. A radiant head is a variant of a discoid head, with peripheral disk flower corollas expanded, often bilateral. A ray flower corolla is bilateral, generally with a slender tube and flattened petal-like ray (single lip composed of generally 3 lobes). Ray flowers are generally pistillate or sterile (occasionally lacking styles). Radiate heads have peripheral ray flowers and central disk flowers. Disciform heads superficially resemble discoid heads, with pistillate or sterile flowers that lack rays, together with or separate from disk flowers. A ligulate flower is bisexual, with a bilateral, generally ephemeral corolla and 5-lobed ligule. Liguliflorous heads comprise only ligulate flowers. See glossary p. 31 for illustrations of family characteristics. Echinops sphaerocephalus L., Gaillardia aristata Pursh, Gaillardia pulchella Foug., Hymenothrix loomisii S.F. Blake, Tagetes erecta L., Thelesperma megapotamicum (Spreng.) Kuntze are waifs. Melampodium perfoliatum Kunth, historic urban waif. Ageratum conyzoides L., Guizotia abyssinica (L. f.) Cass., Santolina chamaecyparisus L., orth. var. are rare or uncommon escapes from cultivation. Dyssodia papposa, Ismelia carinata (Schousb.) Sch. Bip. [Chrysanthemum carinatum Schousb.], Mantisalca salmantica (L.) Briq. & Cavill. are historical or extirpated waifs in California. Inula helenium L. not documented in California. Taxa of Aster in TJM (1993) treated here in Almutaster, Eucephalus, Eurybia, Ionactis, Oreostemma, Sericocarpus, Symphyotrichum; Chamomilla in Matricaria; Bahia in Hymenothrix; Cnicus in Centaurea; Conyza in Erigeron and Laennecia; Dugaldia in Hymenoxys; Erechtites in Senecio; Hymenoclea in Ambrosia; Lembertia in Monolopia; Osteospermum ecklonis in Dimorphotheca; Picris echioides in Helminthotheca; Prionopsis in Grindelia; Raillardiopsis in Anisocarpus and Carlquistia; Schkuhria multiflora in Picradeniopsis; Trimorpha in Erigeron; Venidium in Arctotis; Whitneya in Arnica. Amauriopsis in TJM2 (2012) treated here in Hymenothrix; Arida in Leucosyris; Bahia in Picradeniopsis.
Unabridged Note: Largest family of vascular plants in California and of eudicots globally.
eFlora Treatment Author: David J. Keil, except as noted
Scientific Editor: David J. Keil, Bruce G. Baldwin.
Genus: AdenophyllumView DescriptionDichotomous Key

Habit: [Annual] perennial herb, subshrub [shrub], 20--50+ cm, pungently scented. Stem: erect, branched from base or throughout. Leaf: alternate to +- opposite [opposite], petioled or sessile; blades or lobes linear to oblanceolate or ovate, simple or 1-pinnately lobed, with +- bristle-tipped teeth, glabrous or +- minutely coarse-hairy to minutely scabrous, embedded oil-glands generally at bases of lobes or blade and near leaf tip. Inflorescence: heads generally radiate, 1, peduncled; bracts closely subtending head 12--22, linear to awl-shaped, sometimes 1--2-lobed, often bristle-tipped, generally with oil-glands; involucre bell-shaped to obconic [hemispheric], [5]7--20+ mm diam; phyllaries [8]12--20+ in +- 2 series, weakly fused 1/3--3/4 lengths, free in age, generally lanceolate to linear, persistent, margins of outer +- free to base, oil-glands on faces and near margins; receptacle convex, epaleate, +- pitted, margins of pits minutely fringed to bristly. Ray Flower: (0)7--14[16]; corolla yellow to orange or red-orange. Disk Flower: 25--80[100+]; corolla yellow to yellow-orange, tube < cylindric throat, lobes lance-ovate to -linear; anther tip elliptic or ovate, acute; style tips linear, tapered, densely bristly.; anther bases minutely sagittate. Fruit: obpyramidal [obconic], glabrous or +- strigose or silky-hairy, pappus of 8--20 scales in 1[2] series, each scale composed of 5--11 basally fused bristles.
Species In Genus: 10 species: southwestern United States, Mexico, Central America. Etymology: (Greek: gland-leaf)
eFlora Treatment Author: Bruce G. Baldwin, adapted from Strother (2006)
Reference: Strother 1986 Sida 11:371--378; Strother 2006 FNANM 21:237--239

Adenophyllum cooperi (A. Gray) Strother
Habit: Plant 25--50+ cm; herbage glaucous. Leaf: blade 8--25 mm, 4--15 mm wide, sometimes +- lobed at base, +- glabrous to densely puberulent, oil-glands 1--2 pairs near base, 1 near tip of each leaf. Inflorescence: peduncle 6--15 cm; bracts closely subtending head 12--22, 5--8 mm, linear, long-tapered, glandular; involucre 15--18 mm, bell-shaped to obconic; phyllaries +- 20, lanceolate to linear. Ray Flower: (0)7--14; corolla yellow-orange, becoming red-orange, tube 5 mm, ray 8--10 mm, 2.5--4 mm wide. Disk Flower: corolla 8--10 mm, yellow. Fruit: 5--7 mm, 3-angled; pappus scales 7--10 mm, each composed of 5--9 basally fused bristles. Chromosomes: 2n=26.
Ecology: Washes, alluvial fans, sometimes rocky slopes, in open scrub, woodland; Elevation: 450--1600 m. Bioregional Distribution: n edge SnBr, DMoj; Distribution Outside California: to southwestern Utah, northwestern Arizona. Flowering Time: Apr--Jun, Sep--Nov
Synonyms: Dyssodia cooperi A. Gray;
Jepson eFlora Author: Bruce G. Baldwin, adapted from Strother (2006)
Reference: Strother 1986 Sida 11:371--378; Strother 2006 FNANM 21:237--239
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botanical illustration including Adenophyllum cooperi


Citation for this treatment: Bruce G. Baldwin, adapted from Strother (2006) 2012, Adenophyllum cooperi, in Jepson Flora Project (eds.) Jepson eFlora,, accessed on November 14, 2018.

Citation for the whole project: Jepson Flora Project (eds.) 2018, Jepson eFlora,, accessed on November 14, 2018.

Adenophyllum cooperi
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© 2010 Aaron Schusteff
Adenophyllum cooperi
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© 1997 Christopher L. Christie
Adenophyllum cooperi
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© 2010 Neal Kramer
Adenophyllum cooperi
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© 2013 Neal Kramer
Adenophyllum cooperi
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© 2013 Neal Kramer
Adenophyllum cooperi
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© 2003 James M. Andre

More photos of Adenophyllum cooperi in CalPhotos

Geographic subdivisions for Adenophyllum cooperi:
n edge SnBr, DMoj;
Markers link to CCH specimen records. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues. Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa, if there are more than 1 infraspecific taxon in CA.
Blue line denotes eFlora flowering time.