Common Name: BORAGE or WATERLEAF FAMILY
Habit: Annual to shrub or small tree, or non-green root parasite, often bristly or sharp-hairy. Stem: prostrate to erect. Leaf: basal and/or cauline, generally simple, generally alternate. Inflorescence: generally cymes, or panicle-, raceme-, head-, or spike-like, generally coiled in flower (often described as scorpioid), generally elongating in fruit, or flowers 1--2 per axil. Flower: bisexual, generally radial; sepals (4)5(10), fused at least at base, or free; corolla (4)5(10)-lobed, salverform, funnel-shaped, rotate, or bell-shaped, generally without scales at tube base, with 0 or 5 appendages at tube top, alternate stamens; stamens epipetalous; ovary generally superior, entire to 4-lobed, style 1(2), entire or 2-lobed or -branched. Fruit: valvate or circumscissile capsule or nutlets 1--4, free (fused), smooth to roughened, prickly or bristly or not.
Genera In Family: +- 120 genera, +- 2300 species: tropics, temperate, especially western North America, Mediterranean; some cultivated (Borago, Heliotropium, Echium, Myosotis, Nemophila, Phacelia, Symphytum, Wigandia). Toxicity: Many genera may be TOXIC from pyrrolizidine alkaloids or accumulated nitrates. Note: Recently treated to include Hydrophyllaceae, Lennoaceae. Wigandia urens added, as naturalized.
eFlora Treatment Author: Ronald B. Kelley, Robert Patterson, Richard R. Halse & Timothy C. Messick, family description, key to genera; treatment of genera by Ronald B. Kelley, except as noted
Scientific Editor: Ronald B. Kelley, Robert Patterson, Thomas J. Rosatti, Bruce G. Baldwin, David J. Keil.
Species In Genus: 1 sp. Etymology: (Greek: abiding flower, from persistent corolla)
eFlora Treatment Author: Robert Patterson & Richard R. Halse
Common Name: WHISPERING BELLS
Habit: Annual, glandular, sticky, odorous. Stem: erect, 5--85 cm, branches 0--many. Leaf: basal and cauline, alternate; lower short-petioled, upper sessile, generally clasping, 1--12 cm, generally < 3 cm wide, toothed to deep-pinnate-lobed. Inflorescence: terminal; pedicels 5--15 mm, 12--25 mm in fruit, thread-like, +- subspreading to recurved. Flower: calyx lobes 4--11 mm, 1--4 mm wide, lanceolate to ovate, glandular; corolla 6--15 mm, bell-shaped, white, yellow, or pink, glandular-hairy, persistent in age, withering, papery, enclosing fruit; stamens included, generally equal, attached +- at same level; ovary chambers appearing 2, style included, 1--4 mm, lobes 2. Fruit: capsule, 7--10 mm, 2--4 mm wide, glandular. Seed: 6--15, flat, wide-elliptic, brown; surface honeycombed. Chromosomes: n=18.