Common Name: BORAGE or WATERLEAF FAMILY
Habit: Annual to shrub or small tree, or non-green root parasite, often bristly or sharp-hairy. Stem: prostrate to erect. Leaf: basal and/or cauline, generally simple, generally alternate. Inflorescence: generally cymes, or panicle-, raceme-, head-, or spike-like, generally coiled in flower (often described as scorpioid), generally elongating in fruit, or flowers 1--2 per axil. Flower: bisexual, generally radial; sepals (4)5(10), fused at least at base, or free; corolla (4)5(10)-lobed, salverform, funnel-shaped, rotate, or bell-shaped, generally without scales at tube base, with 0 or 5 appendages at tube top, alternate stamens; stamens epipetalous; ovary generally superior, entire to 4-lobed, style 1(2), entire or 2-lobed or -branched. Fruit: valvate or circumscissile capsule or nutlets 1--4, free (fused), smooth to roughened, prickly or bristly or not.
Genera In Family: +- 120 genera, +- 2300 species: tropics, temperate, especially western North America, Mediterranean; some cultivated (Borago, Heliotropium, Echium, Myosotis, Nemophila, Phacelia, Symphytum, Wigandia). Toxicity: Many genera may be TOXIC from pyrrolizidine alkaloids or accumulated nitrates. Note: Recently treated to include Hydrophyllaceae, Lennoaceae. Wigandia urens added, as naturalized.
eFlora Treatment Author: Ronald B. Kelley, Robert Patterson, Richard R. Halse & Timothy C. Messick, family description, key to genera, treatment of genera by Ronald B. Kelley, except as noted
Scientific Editor: Ronald B. Kelley, Robert Patterson, Thomas J. Rosatti, Bruce G. Baldwin, David J. Keil.
Habit: Annual to subshrub, variously hairy, glandular or not; +- taprooted, rhizome generally 0. Stem: prostrate. Leaf: cauline, generally clustered, evergreen, petioled, margin rolled under, entire or +- crenate. Inflorescence: +- axillary; flowers 1 or clustered, sessile. Flower: radial to +- bilateral; calyx lobes 5, not enlarged in fruit; corolla generally +- funnel-shaped, tube yellow in youth, appendages 0; style 2-lobed, stigmas 2. Fruit: nutlets 1--4, not separate to base, 4-grooved to deep-4-lobed, +- tubercled or not.
Species In Genus: 27 species: western hemisphere deserts. Etymology: (Native South American name for flower)
Unabridged Note: Separated from Coldenia of eastern hemisphere prior to TJM (1993).
eFlora Treatment Author: Ronald B. Kelley
Common Name: SHRUBBY TIQUILIA
Habit: Perennial herb, subshrub, woody. Stem: branches alternate, many, hairs +- spreading. Leaf: occasionally clustered, white-tomentose; blade 5--13 mm, ovate to narrow-elliptic, veins obscure, margin entire, spiny-ciliate. Inflorescence: flowers 1 or clustered in leaf axils; bracts 0. Flower: calyx 3--5(6) mm, lobes 2/3--3/4 free; style short-exserted from calyx, branched < 1/3. Fruit: spheric, 4-grooved, not lobed; nutlets 2--2.5 mm, ovoid, minutely tubercled, hairy or not.