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Plagiobothrys collinus var. gracilis
SAN DIEGO POPCORNFLOWER

Higher Taxonomy
Family: BoraginaceaeView DescriptionDichotomous Key

Common Name: BORAGE or WATERLEAF FAMILY
Habit: Annual to shrub or small tree, or non-green root parasite, often bristly or sharp-hairy. Stem: prostrate to erect. Leaf: basal and/or cauline, generally simple, generally alternate. Inflorescence: generally cymes, or panicle-, raceme-, head-, or spike-like, generally coiled in flower (often described as scorpioid), generally elongating in fruit, or flowers 1--2 per axil. Flower: bisexual, generally radial; sepals (4)5(10), fused at least at base, or free; corolla (4)5(10)-lobed, salverform, funnel-shaped, rotate, or bell-shaped, generally without scales at tube base, with 0 or 5 appendages at tube top, alternate stamens; stamens epipetalous; ovary generally superior, entire to 4-lobed, style 1(2), entire or 2-lobed or -branched. Fruit: valvate or circumscissile capsule or nutlets 1--4, free (fused), smooth to roughened, prickly or bristly or not.
Genera In Family: +- 120 genera, +- 2300 species: tropics, temperate, especially western North America, Mediterranean; some cultivated (Borago, Heliotropium, Echium, Myosotis, Nemophila, Phacelia, Symphytum, Wigandia). Toxicity: Many genera may be TOXIC from pyrrolizidine alkaloids or accumulated nitrates. Note: Recently treated to include Hydrophyllaceae, Lennoaceae. Wigandia urens added, as naturalized.
eFlora Treatment Author: Ronald B. Kelley, Robert Patterson, Richard R. Halse & Timothy C. Messick, family description, key to genera; treatment of genera by Ronald B. Kelley, except as noted
Scientific Editor: Ronald B. Kelley, Robert Patterson, Thomas J. Rosatti, Bruce G. Baldwin, David J. Keil.
Genus: PlagiobothrysView DescriptionDichotomous Key

Common Name: POPCORNFLOWER
Habit: Annual (perennial herb), generally strigose to spreading-hairy; fibrous- to taprooted, staining red dye present or not. Stem: branched at base or above, < 5 dm. Leaf: cauline or basal and cauline, 0.5--10 cm, generally smaller tipward, linear to oblanceolate. Inflorescence: raceme- or spike-like cymes, coiled in bud, generally elongate in fruit; bracts 0--many. Flower: calyx lobes fused below middle, 2--10 mm in fruit; corolla rotate to funnel-shaped or cylindric, white or white with yellow area, tube generally +- yellow inside, limb 1--12 mm diam, appendages prominent to minute, white to yellow. Fruit: nutlets generally 4, +- ovate (triangular to +- lanceolate), rarely on narrow stalk or short peg, variously roughened, abaxially generally with central ridge, lateral ridges, cross-ribs, generally tubercled, occasionally prickly or bristly; adaxially keeled above attachment scar, scar on side generally near middle to base, sometimes on bottom or oblique (on angle between side and bottom), generally raised.
Species In Genus: +- 65 species: temperate western North America, western South America, northeastern Asia, Australia. Etymology: (Greek: sideways pit, from position of nutlet attachment scar) Note: Nutlet characters in key generally best for 3 nutlets farthest from stem; yellow on corolla changes to white after pollination.
Unabridged Note: Fully mature nutlets needed for identification; in many species nutlet closest to stem often more firmly attached, larger, differently textured, and with completely different attachment scar than other 3; nutlet characters used in key focus on 3 more loosely attached nutlets. Intergradation common in some species groups; reticulate speciation in genus; sect. Allocarya often treated as separate genus; many species need study. Corolla size can diminish markedly during flower period. Yellow corolla appendages and, if present, contrasting yellow corolla centers, change to white after successful pollination.
Species: Plagiobothrys collinusView Description 

Habit: Annual, hairs +- spreading, fine or coarse. Stem: prostrate to erect, generally 0.2--4 dm. Leaf: cauline, lower generally opposite, 1--4 cm, upper alternate. Inflorescence: generally elongate, bracts at least below; pedicel to 1.5 mm. Flower: calyx +- 3 mm; corolla limb 1--8 mm diam, appendages generally inconspicuous, +- white. Fruit: nutlet 1--2 mm, ovate; abaxial ridge, lateral ridges, cross-ribs sharp; scar near middle, at tip of +- 0.5 mm, narrow, angled stalk.

Unabridged Note: Typical var. in Chile. Like many of the chaparral- and scrub-associated Plagiobothrys species, often appears after fires. Plagiobothrys collinus var. californicus, Plagiobothrys collinus var. fulvescens, Plagiobothrys collinus var. gracilis +- intergrade.

Plagiobothrys collinus var. gracilis (I.M. Johnst.) Higgins
NATIVE
Habit: Plant not cespitose. Stem: +- prostrate, 1--3 dm, hairs generally coarse. Leaf: 1--4 cm, 2--2.5 mm wide, linear. Inflorescence: elongate, exceeding leaves. Flower: corolla limb 1.5--2 mm diam. Fruit: nutlet 1--1.5 mm.
Ecology: Dry places, +- clay openings in coastal scrub, occasionally chaparral; Elevation: 5--640 m. Bioregional Distribution: SCo (Riverside, San Diego cos.), ChI, sw PR (w San Diego Co.). Flowering Time: Jan--May
Synonyms: Plagiobothrys californicus (A. Gray) Greene var. gracilis I.M. Johnst.
eFlora Treatment Author: Ronald B. Kelley, Robert Patterson, Richard R. Halse & Timothy C. Messick, family description, key to genera; treatment of genera by Ronald B. Kelley
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Citation for this treatment: Ronald B. Kelley, Robert Patterson, Richard R. Halse & Timothy C. Messick, family description, key to genera; treatment of genera by Ronald B. Kelley 2016. Plagiobothrys collinus var. gracilis, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=63890, accessed on February 11, 2016.

Citation for the whole project: Jepson Flora Project (eds.) 2016. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on February 11, 2016.


Plagiobothrys collinus var. gracilis
click for enlargement
© 2014 Steve Matson
Plagiobothrys collinus var. gracilis
click for enlargement
© 2014 Steve Matson
Plagiobothrys collinus var. gracilis
click for enlargement
© 2014 Steve Matson
Plagiobothrys collinus var. gracilis
click for enlargement
© 2014 Steve Matson
Plagiobothrys collinus var. gracilis
click for enlargement
© 2014 Steve Matson

More photos of Plagiobothrys collinus var. gracilis in CalPhotos


Geographic subdivisions for Plagiobothrys collinus var. gracilis:
SCo (Riverside, San Diego cos.), ChI, sw PR (w San Diego Co.).
Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.