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Vascular Plants of California
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Guilleminea densa var. aggregata

Higher Taxonomy
Family: AmaranthaceaeView DescriptionDichotomous Key
Habit: Annual to subshrub; monoecious and/or dioecious; occasionally spiny; hairs simple (branched). Leaf: blade simple, alternate or opposite, margins entire or serrate; veins pinnate; stipules 0. Inflorescence: axillary or terminal; 3-flowered cymes in dense spikes, heads or panicles; bracts 0 or 1--5, persistent; bractlets 0--2. Flower: bisexual or unisexual, small, green (+- white), yellow or purple; perianth parts 0 or (1)3--5, free or fused basally, scarious or hardened, persistent; stamens 1--5, opposite perianth parts, free or basally fused as a tube, generally unequal, occasionally alternate with appendages on stamen tubes (pseudostaminodes), anthers 2- or 4-chambered; ovary superior, chamber 1; ovule 1 (2--many); style (0)1--3, stigmas 1--3(5). Fruit: utricle; generally with persistent perianth or bracts. Seed: 1 [2+], small, lenticular to spheric, smooth or dotted to striate or tubercled.
Genera In Family: +- 75 genera, 900 species: cosmopolitan, especially disturbed, arid, saline or alkaline soils; some cultivated for food, ornamental; many naturalized, ruderal or agricultural weeds. Note: Amaranthaceae including Chenopodiaceae by some. Polycnemoideae, represented in California by Nitrophila, formerly considered subfamily of Chenopodiaceae, but needs further research. Guilleminea densa (Willd.) Moq. var. aggregata Uline & W.L. Bray is a waif. Froelichia gracilis (Hook.) Moq. possibly naturalized.
eFlora Treatment Author: Mihai Costea, except as noted
Scientific Editor: Douglas H. Goldman, Bruce G. Baldwin.
Genus: GuillemineaView Description 

Habit: Perennial herb, generally prostrate, soft-hairy to woolly, hairs simple. Leaf: opposite; petioled; blade linear or lanceolate to spoon-shaped or ovate, margins entire. Inflorescence: axillary, sessile, few-flowered heads; bracts, bractlets membranous, white, glabrous, persistent. Flower: bisexual; perianth bell-shaped, perianth parts 5, fused +- 1/2, 1-veined; stamens 5, filaments fused basally into tube attached to top of perianth tube, anthers 2-chambered, pseudostaminodes 0; ovary ovoid, ovule 1, style 1, +- 0.2 mm, stigmas 2-lobed, spheric. Fruit: utricle, wall membranous. Seed: 1 lenticular to spheric, smooth, shiny.
Species In Genus: +- 5 species: southern United States to South America. Etymology: (Antoine Guillemin, French botanist, explorer, 1796--1842)
Reference: Henrickson 1987 Sida 12:307--337
Guilleminea densa (Willd.) Moq. var. aggregata Uline & W.L. Bray
Habit: Mats 10--80 cm wide. Stem: +- densely woolly. Leaf: basal in rosettes, withering, petiole winged, 8--20 mm, blade(8)20--40 mm, (1)6--9(13) mm wide, oblong-elliptic to lanceolate, glabrous adaxially, woolly abaxially; cauline petiole 1--5 mm, blade 20--100(220) mm, 20--80(150) mm wide, ovate, oblong-ovate, oblong, rhombic, or occasionally elliptic to lanceolate, +- glabrous adaxially, woolly abaxially. Inflorescence: axillary clusters of heads 2--3 mm, 3--5 mm wide, white; bracts 1.5--2.5 mm, white-scarious. Flower: perianth 0.8--2.8 mm, tube 0.8--1.2 mm, lobes reflexed, erect in age, ovate to elliptic, tip acute to rounded, shaggy-hairy especially near tube base. Fruit: included within persistent perianth, 1.1--1.4 mm. Seed: 0.8--0.9 mm, red-brown.
Ecology: Roadsides, disturbed areas; Elevation: < 100 m. Bioregional Distribution: s CCo, SCo; Distribution Outside California: native Arizona to Texas, South America. Flowering Time: Aug--Nov
Unabridged Synonyms: Brayulinea densa (Willd.) Small
Jepson eFlora Author: Mihai Costea
Reference: Henrickson 1987 Sida 12:307--337
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Citation for this treatment: Mihai Costea 2012, Guilleminea densa var. aggregata, in Jepson Flora Project (eds.) Jepson eFlora,, accessed on August 18, 2019.

Citation for the whole project: Jepson Flora Project (eds.) 2019, Jepson eFlora,, accessed on August 18, 2019.

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Geographic subdivisions for Guilleminea densa var. aggregata:
s CCo, SCo
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Data provided by the participants of the  Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa, if there are more than 1 infraspecific taxon in CA.
Blue line denotes eFlora flowering time.