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Vascular Plants of California
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Athyrium filix-femina var. cyclosorum


Higher Taxonomy
Family: AthyriaceaeView Description 
Common Name: LADY FERN FAMILY
Habit: Perennial herb in soil [on rocks]; rhizome short- to long-creeping or tree-like. Leaf: generally tufted or short-spaced, 5--200+ cm, generally +- alike; stipe firm or fleshy (easily crushed), base darker or not, with 2 vascular strands, often expanded at base ("trophopods"); blade generally 1--3-pinnate, +- glabrous or with hairs, hair-like scales, or gland-tipped hairs on axes, veins generally free (or netted); rachis, costa generally grooved adaxially. Sporangia: sori linear along veins, sometimes appearing round; indusia 0 or linear or J-shaped (round), opening away from the vein, when linear, generally back-to-back along a vein.
Genera In Family: 3--5 genera, +- 600 species: worldwide, especially tropics, wooded areas. Note: Comprises three diverse genera (mostly temperate Athyrium, Deparia; mostly tropical Diplazium), and smaller genera often segregated from Athyrium (Anisocampium, Cornopteris; Rothfels et al. 2012 Taxon 61:515--530). One of the most diverse families of the eupolypods II clade (suborder Aspleniineae; Pteridophyte Phylogeny Group 1 2016 J Syst Evol in press); most closely related to Blechnaceae and Onocleaceae (Rothfels et al. 2012 Syst Biol 61:490--509), and more distantly to Woodsiaceae, where often included (Smith et al. 2016 Taxon 55:705--731).
eFlora Treatment Author: Carl J. Rothfels & Alan R. Smith
Scientific Editor: Bruce G. Baldwin & Thomas J. Rosatti.
Genus: AthyriumView DescriptionDichotomous Key


Common Name: LADY FERN
Habit: Rhizome short-creeping to suberect, stout. Leaf: stipe stout, fleshy, easily crushed, straw-colored except base generally blackened, base scaly, ×-section with 2 crescent-shaped vascular strands; blade generally >= 2-pinnate, pinnae of equal sides, +- glabrous or minutely hairy, veins free. Sporangia: sori +- round, +- oblong, or J-shaped; indusia 0, oblong, J-shaped, or reniform, laterally attached.
Species In Genus: +- 100 species: generally northern temperate, especially eastern Asia. Etymology: (Greek: doorless, from enclosed sori)
eFlora Treatment Author: Carl J. Rothfels & Alan R. Smith
Athyrium filix-femina (L.) Roth var. cyclosorum Rupr.
NATIVE
Leaf: blade elliptic to lanceolate, 1--2-pinnate-pinnatifid, lower 2--4 pinna pairs generally << those above, ultimate segments pinnately lobed to +- toothed, midribs below near base often with minute, branched hairs 0.1--0.2 mm. Chromosomes: 2n=80.
Ecology: Woodland, along streams, seepage areas; Elevation: < 3200 m. Bioregional Distribution: CA-FP (exc Teh, ScV, SCoRI, SCo, WTR, PR exc SnJt), MP; Distribution Outside California: to Alaska, western Canada, Idaho, Colorado, northern Central America. Note: Highly variable, but named varieties in western North America seem indistinct; other varieties worldwide.
Synonyms: Athyrium filix-femina var. californicum Butters; Athyrium filix-femina var. sitchense Rupr.
Jepson eFlora Author: Carl J. Rothfels & Alan R. Smith
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Botanical illustration including Athyrium filix-femina var. cyclosorum

botanical illustration including Athyrium filix-femina var. cyclosorum

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Citation for this treatment: Carl J. Rothfels & Alan R. Smith 2016, Athyrium filix-femina var. cyclosorum, in Jepson Flora Project (eds.) Jepson eFlora, Revision 4, http://ucjeps.berkeley.edu/eflora/eflora_display.php?tid=54990, accessed on July 24, 2019.

Citation for the whole project: Jepson Flora Project (eds.) 2019, Jepson eFlora, http://ucjeps.berkeley.edu/eflora/, accessed on July 24, 2019.

Athyrium filix-femina var. cyclosorum
click for enlargement
© 2009 Keir Morse
Athyrium filix-femina var. cyclosorum
click for enlargement
© 2018 California Academy of Sciences
Athyrium filix-femina var. cyclosorum
click for enlargement
© 2008 Keir Morse
Athyrium filix-femina var. cyclosorum
click for enlargement
© 2015 Barry Breckling
Athyrium filix-femina var. cyclosorum
click for enlargement
© 2018 California Academy of Sciences
Athyrium filix-femina var. cyclosorum
click for enlargement
© 2008 Keir Morse

More photos of Athyrium filix-femina var. cyclosorum in CalPhotos



Geographic subdivisions for Athyrium filix-femina var. cyclosorum:
CA-FP (exc Teh, ScV, SCoRI, SCo, WTR, PR exc SnJt), MP
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map of distribution 1
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Duplicates counted once; synonyms included.
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