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Athyrium filix-femina var. cyclosorum

Higher Taxonomy
Family: WoodsiaceaeView DescriptionDichotomous Key

Habit: Perennial herb in soil or rock crevices; rhizome generally short-creeping, ascending, or erect, scales small to large, generally tan to brown, generally uniformly colored. Leaf: generally tufted or short-spaced, 5--200+ cm, generally +- alike; stipe firm or fleshy (easily crushed), base darker or not, with 2 vascular strands; blade generally 1--3-pinnate, +- glabrous or with hairs, hair-like scales, or gland-tipped hairs on axes, veins generally free (or netted); rachis, costa generally grooved adaxially. Sporangia: sori round, oblong, J-shaped, or linear along veins; indusia 0 or oblong, J-shaped, reniform, or linear, or of many segmented hair- or scale-like fragments or lobes encircling sorus from below; spores elliptic, winged, ridged, or spiny, scar linear.
Genera In Family: +- 15 genera, 700 species: worldwide, especially tropics, wooded areas, but some genera (e.g., Cystopteris, Woodsia) generally temperate. Note: See note, reference (Smith et al. 2006 Taxon 55:705--731) under Dryopteridaceae for removal of genera from that family to this.
eFlora Treatment Author: Alan R. Smith, except as noted
Scientific Editor: Thomas J. Rosatti.
Genus: AthyriumView DescriptionDichotomous Key

Common Name: LADY FERN
Habit: Rhizome short-creeping to suberect, stout. Leaf: stipe stout, fleshy, easily crushed, straw-colored except base generally blackened, base scaly, ×-section with 2 crescent-shaped vascular strands; blade generally >= 2-pinnate, pinnae of equal sides, +- glabrous or minutely hairy, veins free. Sporangia: sori +- round, +- oblong, or J-shaped; indusia 0, oblong, J-shaped, or reniform, laterally attached.
Species In Genus: +- 100 species: generally northern temperate, especially eastern Asia. Etymology: (Greek: doorless, from enclosed sori)

Athyrium filix-femina (L.) Roth var. cyclosorum Rupr.
Leaf: blade elliptic to lanceolate, 1--2-pinnate-pinnatifid, lower 2--4 pinna pairs generally << those above, ultimate segments pinnately lobed to +- toothed, midribs below near base often with minute, branched hairs 0.1--0.2 mm. Chromosomes: 2n=80.
Ecology: Woodland, along streams, seepage areas; Elevation: < 3200 m. Bioregional Distribution: CA-FP (exc Teh, ScV, SCoRI, SCo, WTR, PR exc SnJt), MP; Distribution Outside California: to Alaska, western Canada, Idaho, Colorado, northern Central America. Note: Highly variable, but named varieties in western North America seem indistinct; other varieties worldwide.
Synonyms: Athyrium filix-femina var. californicum Butters; Athyrium filix-femina var. sitchense Rupr.
eFlora Treatment Author: Alan R. Smith
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Previous taxon: Athyrium distentifolium var. americanum
Next taxon: Cystopteris

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botanical illustration including Athyrium filix-femina var. cyclosorum


Citation for this treatment: Alan R. Smith 2016. Athyrium filix-femina var. cyclosorum, in Jepson Flora Project (eds.) Jepson eFlora,, accessed on May 26, 2016.

Citation for the whole project: Jepson Flora Project (eds.) 2016. Jepson eFlora,, accessed on May 26, 2016.

Athyrium filix-femina var. cyclosorum
click for enlargement
© 2015 Barry Breckling
Athyrium filix-femina var. cyclosorum
click for enlargement
© 2009 Keir Morse
Athyrium filix-femina var. cyclosorum
click for enlargement
© 2008 Keir Morse
Athyrium filix-femina var. cyclosorum
click for enlargement
© 2008 Keir Morse
Athyrium filix-femina var. cyclosorum
click for enlargement
© 2008 Keir Morse
Athyrium filix-femina var. cyclosorum
click for enlargement
© 2008 Keir Morse

More photos of Athyrium filix-femina var. cyclosorum in CalPhotos

Geographic subdivisions for Athyrium filix-femina var. cyclosorum:
CA-FP (exc Teh, ScV, SCoRI, SCo, WTR, PR exc SnJt), MP;
Markers link to CCH specimen records. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues. Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.