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Oenothera elata subsp. hirsutissima

Higher Taxonomy
Family: OnagraceaeView DescriptionDichotomous Key

Habit: Annual to perennial herb (to tree). Leaf: cauline or basal, alternate, opposite, or whorled, generally simple and toothed (to pinnately compound); stipules 0 or generally deciduous. Inflorescence: spike, raceme, panicle, or flowers 1 in axils; bracted. Flower: generally bisexual, generally radial, often opening at either dawn or dusk; hypanthium generally prolonged beyond ovary (measured from ovary tip to sepal base); sepals 4(2--7); petals 4(2--7, rarely 0), often fading darker; stamens 2 × or = sepals in number, anthers 2-chambered, opening lengthwise, pollen interconnected by threads; ovary inferior, chambers generally as many as sepals (sometimes becoming 1), placentas axile or parietal, ovules 1--many per chamber, style 1, stigma 4-lobed (or lobes as many as sepals), club-shaped, spheric, or hemispheric. Fruit: capsule, loculicidal (sometimes berry or indehiscent and nut-like). Seed: sometimes winged or hair-tufted.
Genera In Family: 22 genera, +- 657 species: worldwide, especially western North America; many cultivated (Clarkia, Epilobium, Fuchsia, Oenothera). Note: Gaura moved to Oenothera. Fuchsia magellanica Lam. naturalized in northern California.
eFlora Treatment Author: Warren L. Wagner & Peter C. Hoch, family description, key to genera, treatment of genera by Warren L. Wagner, except as noted
Scientific Editor: Robert Patterson, Bruce G. Baldwin.
Genus: OenotheraView DescriptionDichotomous Key

Habit: Annual to perennial herb, generally from taproot, occasionally rhizomed. Leaf: basal or cauline, alternate, generally pinnately toothed to lobed, generally sessile. Inflorescence: spike, raceme-like, or flowers in axils of distal, reduced leaves. Flower: radial or (sect. Gaura) bilateral, generally opening at dusk; sepals 4, reflexed in flower (sometimes 2--3 remaining adherent); petals 4, yellow, white, rose, or +- purple, generally fading +- orange to +- purple, tip notched or toothed; stamens 8, filaments sometimes (sect. Gaura) with paired teeth at base, anthers attached at middle; ovary chambers 4, stigma generally deeply lobed, generally > anthers and cross-pollinated (or +- = anthers and self-pollinated). Fruit: generally dehiscent, cylindric to ovoid or obovoid, cylindric to 4-winged or -angled, straight to curved, generally sessile (base sometimes seedless, stalk-like). Seed: in generally 2(1--3) rows per chamber, or clustered or reduced to 1--4 per fruit.
Species In Genus: 145 species: America, some widely naturalized. Etymology: (Greek: wine-scented)
Unabridged Note: Many species self-pollinated; some of these have chromosome peculiarities (ring of 14 in meiosis) and +- 50% pollen fertility; they yield genetically identical offspring.
Species: Oenothera elataView Description 

Habit: Biennial, densely minutely strigose and (especially in inflorescence) glandular; hairs also long, appressed to spreading, sometimes with red, blister-like base. Stem: erect. Leaf: cauline 4--25 cm, oblanceolate to lanceolate or elliptic, generally dentate to +- entire. Inflorescence: spike. Flower: hypanthium 20--48(55) mm; sepals 27--48 mm; petals 25--52 mm, yellow fading red-orange. Fruit: 20--65 mm, 4--7 mm wide, narrowly lanceolate, +- straight. Seed: 1--1.8 mm, angled, irregularly pitted. Chromosomes: 2n=14.
Note: Generally cross-pollinated. 3 subspecies, 2 in California.

Oenothera elata subsp. hirsutissima (S. Watson) W. Dietr.
Stem: 10--25 dm. Flower: sepals green or red-flushed, hairs glandular or not, also spreading, red, blister-like base +- 0, free tips in bud 3--6 mm; anthers 8--15 mm.
Ecology: Moist places, generally inland; Elevation: < 2800 m. Bioregional Distribution: CA; Distribution Outside California: western United States, northwestern Mexico. Flowering Time: Jun--Sep Note: Several intergrading forms.
Synonyms: Oenothera hookeri Torr. & A. Gray subsp. angustifolia (R.R. Gates) Munz; Oenothera hookeri subsp. grisea (Bartlett) Munz; Oenothera hookeri subsp. venusta (Bartlett) Munz
eFlora Treatment Author: Warren L. Wagner & Peter C. Hoch, family description, key to genera, treatment of genera by Warren L. Wagner
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botanical illustration including Oenothera elata subsp. hirsutissima


Citation for this treatment: Warren L. Wagner & Peter C. Hoch, family description, key to genera, treatment of genera by Warren L. Wagner 2017. Oenothera elata subsp. hirsutissima, in Jepson Flora Project (eds.) Jepson eFlora,, accessed on June 24, 2017.

Citation for the whole project: Jepson Flora Project (eds.) 2017. Jepson eFlora,, accessed on June 24, 2017.

Oenothera elata subsp. hirsutissima
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© 2012 Steve Matson
Oenothera elata subsp. hirsutissima
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© 2003 Michael Charters
Oenothera elata subsp. hirsutissima
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© 2009 Thomas Stoughton
Oenothera elata subsp. hirsutissima
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© 2006 George W. Hartwell
Oenothera elata subsp. hirsutissima
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© 2012 Neal Kramer
Oenothera elata subsp. hirsutissima
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© 2011 Neal Kramer

More photos of Oenothera elata subsp. hirsutissima in CalPhotos

Geographic subdivisions for Oenothera elata subsp. hirsutissima:
Markers link to CCH specimen records. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues. Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.