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Oenothera deltoides subsp. piperi

Higher Taxonomy
Family: OnagraceaeView DescriptionDichotomous Key

Habit: Annual to perennial herb (to tree). Leaf: cauline or basal, alternate, opposite, or whorled, generally simple and toothed (to pinnately compound); stipules 0 or generally deciduous. Inflorescence: spike, raceme, panicle, or flowers 1 in axils; bracted. Flower: generally bisexual, generally radial, often opening at either dawn or dusk; hypanthium generally prolonged beyond ovary (measured from ovary tip to sepal base); sepals 4(2--7); petals 4(2--7, rarely 0), often fading darker; stamens 2 × or = sepals in number, anthers 2-chambered, opening lengthwise, pollen interconnected by threads; ovary inferior, chambers generally as many as sepals (sometimes becoming 1), placentas axile or parietal, ovules 1--many per chamber, style 1, stigma 4-lobed (or lobes as many as sepals), club-shaped, spheric, or hemispheric. Fruit: capsule, loculicidal (sometimes berry or indehiscent and nut-like). Seed: sometimes winged or hair-tufted.
Genera In Family: 22 genera, +- 657 species: worldwide, especially western North America; many cultivated (Clarkia, Epilobium, Fuchsia, Oenothera). Note: Gaura moved to Oenothera. Fuchsia magellanica Lam. naturalized in northern California.
eFlora Treatment Author: Warren L. Wagner & Peter C. Hoch, family description, key to genera; treatment of genera by Warren L. Wagner, except as noted
Scientific Editor: Robert Patterson, Bruce G. Baldwin.
Genus: OenotheraView DescriptionDichotomous Key

Habit: Annual to perennial herb, generally from taproot, occasionally rhizomed. Leaf: basal or cauline, alternate, generally pinnately toothed to lobed, generally sessile. Inflorescence: spike, raceme-like, or flowers in axils of distal, reduced leaves. Flower: radial or (sect. Gaura) bilateral, generally opening at dusk; sepals 4, reflexed in flower (sometimes 2--3 remaining adherent); petals 4, yellow, white, rose, or +- purple, generally fading +- orange to +- purple, tip notched or toothed; stamens 8, filaments sometimes (sect. Gaura) with paired teeth at base, anthers attached at middle; ovary chambers 4, stigma generally deeply lobed, generally > anthers and cross-pollinated (or +- = anthers and self-pollinated). Fruit: generally dehiscent, cylindric to ovoid or obovoid, cylindric to 4-winged or -angled, straight to curved, generally sessile (base sometimes seedless, stalk-like). Seed: in generally 2(1--3) rows per chamber, or clustered or reduced to 1--4 per fruit.
Species In Genus: 145 species: America, some widely naturalized. Etymology: (Greek: wine-scented)
Unabridged Note: Many species self-pollinated; some of these have chromosome peculiarities (ring of 14 in meiosis) and +- 50% pollen fertility; they yield genetically identical offspring.
Species: Oenothera deltoidesView Description 

Habit: Loosely rosetted; hairs curly or straight, also sometimes glandular, or 0. Stem: decumbent or erect, (< 1)2--10 dm, stout, spongy, peeling. Leaf: cauline 2--15 cm, +- diamond-shaped-obovate to oblanceolate, +- entire to pinnately lobed. Inflorescence: flowers in distal axils; buds nodding. Flower: hypanthium 20--40 mm; sepals 8--30 mm, free tips in bud 0--1.2 mm; petals white fading pink. Fruit: 20--60(80) mm, cylindric, generally curved, +- twisted. Seed: in 1 row per chamber, 1.5--2 mm, obovate, smooth. Chromosomes: 2n=14.
Note: Generally cross-pollinated. 5 subspecies, 4 in California.

Oenothera deltoides subsp. piperi (Munz) W.M. Klein
Habit: Annual; hairs wavy or curly, 1.5--3 mm distally. Stem: generally < 1 dm, generally simple or few-branched from base. Leaf: distal pinnately lobed. Flower: bud tip bluntly acute; free sepal tips 0--1 mm; petals 15--25(30) mm. Fruit: base 3--5 mm wide.
Ecology: Sand, including dunes, sagebrush scrub; Elevation: 850--1800 m. Bioregional Distribution: MP; Distribution Outside California: central Oregon, western Nevada. Flowering Time: Mar--May
eFlora Treatment Author: Warren L. Wagner & Peter C. Hoch, family description, key to genera; treatment of genera by Warren L. Wagner
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botanical illustration including Oenothera deltoides subsp. piperi


Citation for this treatment: Warren L. Wagner & Peter C. Hoch, family description, key to genera; treatment of genera by Warren L. Wagner 2016. Oenothera deltoides subsp. piperi, in Jepson Flora Project (eds.) Jepson eFlora,, accessed on May 30, 2016.

Citation for the whole project: Jepson Flora Project (eds.) 2016. Jepson eFlora,, accessed on May 30, 2016.

Oenothera deltoides subsp. piperi
click for enlargement
© 2012 Gary A. Monroe
Oenothera deltoides subsp. piperi
click for enlargement
© 2009 Gary A. Monroe
Oenothera deltoides subsp. piperi
click for enlargement
© 2012 Gary A. Monroe

More photos of Oenothera deltoides subsp. piperi in CalPhotos

Geographic subdivisions for Oenothera deltoides subsp. piperi:
Markers link to CCH specimen records. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues. Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.