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Galium ambiguum subsp. siskiyouense

Higher Taxonomy
Family: RubiaceaeView DescriptionDichotomous Key

Habit: Annual to tree, vine. Leaf: generally opposite (whorled), entire; stipules generally fused to stem, adjacent pairs occasionally fused, or occasionally leaf-like and appearing like whorled leaves. Inflorescence: cyme, panicle, spike, cluster, or flower 1, generally terminal and +- axillary. Flower: generally bisexual; calyx +- 4(5)-lobed, occasionally 0 (Galium, Crucianella) or 6 (Sherardia); corolla generally radial, 4(5)-lobed; stamens epipetalous, alternate corolla lobes, generally included; ovary generally inferior, chambers generally 2 or 4, style 1(2). Fruit: drupe, berry, or 2 or 4 nutlets [capsule].
Genera In Family: +- 500 genera, 6000 species: worldwide, especially tropics; many cultivated, including Coffea, coffee; Cinchona, quinine; many ornamental. Note: Diodia teres Walter doubtfully in California.
eFlora Treatment Author: Robert E. Preston & Lauramay T. Dempster, except as noted
Scientific Editor: Douglas H. Goldman, Bruce G. Baldwin.
Genus: GaliumView DescriptionDichotomous Key

Common Name: BEDSTRAW
Habit: Annual, perennial herb, occasionally subshrub, glabrous to hairy, generally scabrous; dioecious, bisexual, or flowers unisexual and bisexual. Stem: 4-angled, occasionally ridged lengthwise. Leaf: generally in whorls of >= 4, including leaf-like stipules. Inflorescence: panicles, axillary clusters (cymes), or occasionally 1 in axils. Flower: bisexual, or unisexual with sterile stamens or pistils; calyx 0; corolla generally rotate, occasionally +- bell-shaped, generally +- green, yellow to white, occasionally pink or red, lobes generally 4; ovary 2-lobed, styles 2, bases +- fused. Fruit: 2 nutlets or berry.
Species In Genus: +- 650 species: worldwide, especially temperate. Etymology: (Greek: milk, from use of some species for curdling) Note: Ovary and fruit generally +- equally hairy on a pl; staminate plants generally identified by vestigial ovaries, pistillate plants generally by vestigial anthers. Galium saxatile L., Galium schultesii Vest, and Galium verum L. are lawn weeds in California.
eFlora Treatment Author: Valerie Soza
Species: Galium ambiguumView Description 

Habit: Perennial herb, low, matted, not woody above ground, generally hairy; dioecious. Stem: erect, 5--16 cm. Leaf: in whorls of 4, 6--16 mm, +- linear, tip +- acute, with long bristle, margin generally ciliate. Staminate Inflorescence: few-flowered clusters. Pistillate Inflorescence: flowers generally 1 in axils. Flower: corolla rotate, +- yellow, +- glabrous to short-fine-ciliate or papillate along margins. Fruit: berry, glabrous to hairy.

Galium ambiguum subsp. siskiyouense (Ferris) Dempster & Stebbins
Habit: Coarse-hairy to glabrous. Leaf: generally thick, glabrous or margins bristly, faces occasionally irregularly or minutely hairy, margins generally convex. Fruit: densely hairy. Chromosomes: 2n=66.
Ecology: Dry, sunny banks and hillsides, open forest, in serpentine or clay loam; Elevation: 160--1750 m. Bioregional Distribution: KR (Del Norte Co.), NCoRO; Distribution Outside California: southern Oregon. Flowering Time: Jun--Jul
Synonyms: Galium ambiguum var. siskiyouense Ferris
eFlora Treatment Author: Valerie Soza
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botanical illustration including Galium ambiguum subsp. siskiyouense


Citation for this treatment: Valerie Soza 2017. Galium ambiguum subsp. siskiyouense, in Jepson Flora Project (eds.) Jepson eFlora,, accessed on September 21, 2017.

Citation for the whole project: Jepson Flora Project (eds.) 2017. Jepson eFlora,, accessed on September 21, 2017.

Galium ambiguum subsp. siskiyouense
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© 2016 Keir Morse
Galium ambiguum subsp. siskiyouense
click for enlargement
© 2016 Keir Morse
Galium ambiguum subsp. siskiyouense
click for enlargement
© 2017 Steve Matson
Galium ambiguum subsp. siskiyouense
click for enlargement
© 2013 Dana York
Galium ambiguum subsp. siskiyouense
click for enlargement
© 2017 Steve Matson
Galium ambiguum subsp. siskiyouense
click for enlargement
© 2017 Steve Matson

More photos of Galium ambiguum subsp. siskiyouense in CalPhotos

Geographic subdivisions for Galium ambiguum subsp. siskiyouense:
KR (Del Norte Co.), NCoRO;
Markers link to CCH specimen records. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues. Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.