Common Name: CLIFF FERN FAMILY Habit: Perennial herb in rock crevices, occasionally in soil; rhizome generally short-creeping. Leaf: short-spaced, 5--40 cm, +- alike; stipe firm or fleshy (easily crushed), base darker or not, with 2 vascular strands; blade 1-pinnate to 2-pinnate-pinnately lobed, herbaceous, generally with hairs, hair-like scales, or gland-tipped hairs on axes, veins free, terminating before leaf margin; rachis, costa generally grooved adaxially. Sporangia: sori round; indusia of many segmented hair- or scale-like fragments or lobes encircling sorus from below [sphere-like]. Genera In Family: 1 genus, +- 35 species: worldwide, generally temperate. All species often treated in Woodsia (Shao et al. 2015 PLoS ONE 10; Pteridophyte Phylogeny Group 2016 J Syst Evol in press); four to six segregate genera could be recognized (Schmakov 2015 Turczaninowia 18:11--16). Note: A relatively isolated family within the broad eupolypods II clade (suborder Aspleniineae; Pteridophyte Phylogeny Group 2016 J Syst Evol in press), only distantly related to Cystopteridaceae or Athyriaceae, both often included here (Smith et al. 2016 Taxon 55:705--731). eFlora Treatment Author: Carl J. Rothfels & Alan R. Smith Scientific Editor: Bruce G. Baldwin & Thomas J. Rosatti.
Common Name: CLIFF FERN Habit: Rhizome generally ascending to suberect, short, old stipe bases many. Leaf: often glandular or hairy; stipe base ×-section with 2 vascular strands; blade 1--2-pinnate, segments +- toothed to pinnately lobed, veins free, ending just short of margin. Sporangia: sori round, generally not at margins; indusium cup-like, often of many segmented hair- or scale-like fragments or lobes encircling sorus from below [sphere-like], often of crusty, +- white beads, often obscure in age. Etymology: (J. Woods, Britain, b. 1776) eFlora Treatment Author: John C. Game, Alan R. Smith and Thomas Lemieux Reference: Windham 1993 FNANM 2:270--280 Unabridged Reference: Brown 1964 Beih Nova Hedwigia 16:1--154
Woodsia oregana D.C. Eaton
NATIVE Leaf: 5--25 cm, 1--3.5 cm wide, tip +- acute, unforked; hairs on abaxial leaf axes 0 or +- 0.1 mm, cylindric, non-segmented, glandular; pinnae 0.5--2.5 cm, 0.3--1.3 cm wide, pinnately lobed to 1-pinnate, margin fine-toothed. Sporangia: indusium of segmented hairs. Chromosomes: 2n=76,152. Ecology: Crevices, rock bases; Elevation: 900--2800 m. Bioregional Distribution: KR, CaRH, n&s SNH, SnBr, PR, MP, W&I, DMtns; Distribution Outside California: to British Columbia, eastern Canada, northern United States, Oklahoma, Arizona. Note: Sterile, nonglandular plants distinguished from Cystopteris fragilis by leaf veins that do not quite reach the margin in Woodsia but do in Cystopteris. If recognized taxonomically, plants differing in chromosome number, spore size [Windham 1993], perhaps other ways, assignable to W. oregana subsp. cathcartiana (B.L. Rob.) Windham (2n = 152), W. oregana subsp. oregana (2n = 76); study needed. Unabridged Note: Variable. Like Cystopteris fragilis except glandular hairs generally present in Woodsia, absent in Cystopteris; veins ending just before leaf margin in Woodsia at 10×, at leaf margin in Cystopteris. Jepson eFlora Author: John C. Game, Alan R. Smith and Thomas Lemieux Reference: Windham 1993 FNANM 2:270--280 Index of California Plant Names (ICPN; linked via the Jepson Online Interchange) Previous taxon: Woodsia Next taxon: Woodsia plummerae
Citation for this treatment: John C. Game, Alan R. Smith and Thomas Lemieux 2012, Woodsia oregana, in Jepson Flora Project (eds.) Jepson eFlora, https://ucjeps.berkeley.edu/eflora/eflora_display.php?tid=48612, accessed on April 23, 2024.
Citation for the whole project: Jepson Flora Project (eds.) 2024, Jepson eFlora, https://ucjeps.berkeley.edu/eflora/, accessed on April 23, 2024.
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(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).
Data provided by the participants of the
Consortium of California Herbaria.
MAP LEGEND View all CCH records All markers link to CCH specimen records. The original determination is shown in the popup window.
Blue markers indicate specimens that map to one of the expected Jepson geographic subdivisions (see left map). Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
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CCH collections by month
Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa, if there are more than 1 infraspecific taxon in CA.
Blue line denotes eFlora flowering time (fruiting time in some monocot genera).
