Jepson eFlora: Taxon page
Vascular Plants of California
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Vancouveria hexandra


Higher Taxonomy
Family: BerberidaceaeView DescriptionDichotomous Key
Common Name: BARBERRY FAMILY
Habit: Perennial herb, shrub, [tree], generally rhizomed, caudexed or not, glabrous, glaucous, or hairy. Stem: spreading to erect, branched or not. Leaf: simple, 1--3-ternate, or pinnately compound, basal and cauline, generally alternate, deciduous or evergreen, petioled, stipuled. Inflorescence: generally raceme, spike, or panicle, scapose, terminal, or axillary. Flower: generally bisexual, radial; sepals 6--18 or 0, generally in whorls of 3; petals generally 6, in 2 whorls of 3, or 0; stamens 6--12(13), free or fused at base, in 2 whorls or not, anthers dehiscent by flap-like valves or longitudinal slits; ovary superior, chamber 1, ovules generally 1--10, style 1 or 0, stigma flat or spheric. Fruit: berry, capsule, achene [follicle].
Genera In Family: 16 genera, +- 670 species: temperate, tropics worldwide; some cultivated (Berberis, Epimedium, Nandina (heavenly bamboo), Vancouveria). Note: Lower sepals sometimes called "bracteoles," inner petals "staminodes".
eFlora Treatment Author: Michael P. Williams
Scientific Editor: Thomas J. Rosatti.
Genus: VancouveriaView DescriptionDichotomous Key


Common Name: INSIDE-OUT-FLOWER
Habit: Perennial herb, rhizomes extensive, scales brown. Stem: 0. Leaf: basal, long-petioled, 2--3-ternate; leaflet blades ovate to +- cordate, lobes 3, shallow, teeth 0 or shallow. Inflorescence: raceme or panicle, +- scapose, open, long-peduncled; flowers spreading to pendent. Flower: sepals generally 12--15, 8--9 mm, outer 6--9 << inner 6, bract-like, deciduous, inner petal-like, persistent, in age reflexed; petals 6, < inner sepals, reflexed from base, distally glandular; stamens generally 6, held against ovary, style, anther valves flap-like, pointed tipward; ovules 2--10, style 1, < ovary, persistent, beak-like in fruit, stigma cup-like. Fruit: capsule, 2-valved, generally elliptic. Seed: with oily body for ant dispersal.
Species In Genus: 3 species: temperate western North America. Etymology: (Captain George Vancouver, British explorer, 1757--1798) Note: Pedicel appears to arise from inside flower, from tip instead of base, yielding an upside-down or "inside-out" flower.
Reference: Zhang 2007 Syst Bot 32:81--92
Unabridged Reference: Stearn 1938 J Linn Soc Bot 51:409--535
Vancouveria hexandra (Hook.) C. Morren & Decne.
NATIVE
Leaf: deciduous in fruit, 8--27 cm, adaxially glabrous, abaxially sparse-hairy; petiole generally glabrous, in age straw-colored. Inflorescence: +- raceme, branched below or not; axis, pedicels glabrous. Flower: outer sepals 2--4 mm, inner 5--7 mm; petals 4--6 mm, white, tip strongly reflexed, +- hood-like; filaments red-glandular. Fruit: body 8--10(15) mm, short glandular-hairy. Chromosomes: 2n=12.
Ecology: Conifer forest; Elevation: < 1900 m. Bioregional Distribution: NCo, w KR, n NCoRO; Distribution Outside California: to western Washington. Flowering Time: May--Jul
Jepson eFlora Author: Michael P. Williams
Reference: Zhang 2007 Syst Bot 32:81--92
Unabridged Reference: Stearn 1938 J Linn Soc Bot 51:409--535
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Botanical illustration including Vancouveria hexandra

botanical illustration including Vancouveria hexandra

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Citation for this treatment: Michael P. Williams 2012, Vancouveria hexandra, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/eflora/eflora_display.php?tid=47782, accessed on June 26, 2019.

Citation for the whole project: Jepson Flora Project (eds.) 2019, Jepson eFlora, http://ucjeps.berkeley.edu/eflora/, accessed on June 26, 2019.

Vancouveria hexandra
click for enlargement
© 2016 Neal Kramer
Vancouveria hexandra
click for enlargement
© 2016 Steve Matson
Vancouveria hexandra
click for enlargement
© 2016 Neal Kramer
Vancouveria hexandra
click for enlargement
© 2016 Steve Matson
Vancouveria hexandra
click for enlargement
© 2008 Keir Morse
Vancouveria hexandra
click for enlargement
© 2010 Aaron Schusteff

More photos of Vancouveria hexandra in CalPhotos



Geographic subdivisions for Vancouveria hexandra:
NCo, w KR, n NCoRO
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).





 

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All markers link to CCH specimen records. The original determination is shown in the popup window.
Blue markers indicate specimens that map to one of the expected Jepson geographic subdivisions (see left map). Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa, if there are more than 1 infraspecific taxon in CA.
Blue line denotes eFlora flowering time.