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Utricularia intermedia
FLAT-LEAVED BLADDERWORT

Higher Taxonomy
Family: LentibulariaceaeView DescriptionDichotomous Key

Common Name: BLADDERWORT FAMILY
Habit: Annual, perennial herb, carnivorous, of moist or aquatic habitats. Stem: caudex or stolon, then often with thread-like branches. Leaf: simple, in rosette, or simple or dissected, emerging from caudex or stolon, with minute, carnivorous bladders +- throughout. Inflorescence: raceme or 1-flowered, scapose. Flower: bisexual; calyx lips 2[4], upper 3-lobed, lower 2-lobed, or lips unlobed; corolla 2-lipped, spurred at base, lower lip flat or arched upward, blocking throat or not; stamens 2, epipetalous; ovary superior, chamber 1, placenta generally free-central; stigma unequally 2-lobed, +- sessile. Fruit: capsule, round, 2-valved, circumscissile, or irregularly dehiscent. Seed: generally many, small.
Genera In Family: 3 genera, 330 species: worldwide, especially tropics.
eFlora Treatment Author: Barry A. Rice
Scientific Editor: Thomas J. Rosatti.
Genus: UtriculariaView DescriptionDichotomous Key

Common Name: BLADDERWORT
Habit: Carnivorous by bladders (here treated as modified leaves), into which small organisms are sucked when hairs at opening are triggered [epiphytic]. Stem: submersed or subterranean shoots [rarely caudex]; some aquatic species produce 2 kinds of stems, green (in water or at surface; leaves with flattened or thread-like segments; bladders 0--few) and white (generally buried in substrate; leaves 0; bladders many), the latter not always present in poor collections. Leaf: simple or generally dissected into narrow segments, alternate on stolon, margins often with bristles (visible at 10--30×). Inflorescence: raceme or 1-flowered, emergent; scape slender or wiry, bracts present. Flower: calyx lips 2[4], unlobed; corolla yellow [or not], with red-brown streaks or not, upper lip +- entire, lower entire or 3-lobed, spurred; rarely cleistogamous. Fruit: capsule.
Species In Genus: +- 220 species: worldwide, especially tropics, Australia. Etymology: (Latin: little bag, from bladders) Note: Size variable, often unreliable for identification; distinction between stems, leaves uncertain. Glands inside bladders consist of 2 pairs of oppositely directed arms, angles of divergence between which used (at 150× or more) to identify fresh or pressed specimens.

Utricularia intermedia Hayne
NATIVE
Stem: of 2 kinds, some freely floating, green, with leaves, rarely with bladders, others rooted in mud, white, without leaves, with bladders; winter buds bristly. Leaf: 3--15 mm, 3-parted at base, variously dissected above; ultimate segments < 20, +- linear, flattened, margins with 5--12(20) bristles, +- entire, tip obtuse. Inflorescence: 1--5-flowered; peduncle < 25 cm, < 1 mm diam; pedicel erect in fruit. Flower: corolla 10--16 mm; lower lip +- 2 × upper, +- > or +- = cylindric, acute spur. Fruit: circumscissile. Seed: not seen. Chromosomes: 2n=44.
Ecology: Shallow (< 1 m) water; Elevation: 1200--2700 m. Bioregional Distribution: CaR, SNH, MP; Distribution Outside California: northern United States, Canada; circumboreal. Flowering Time: Jul--Sep Note: In some populations, divergence of bladder gland arm-pairs resembling that of Utricularia ochroleuca.
eFlora Treatment Author: Barry A. Rice
Jepson Online Interchange
Listed on CNPS Rare Plant Inventory

Previous taxon: Utricularia gibba
Next taxon: Utricularia macrorhiza

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Citation for this treatment: Barry A. Rice 2016. Utricularia intermedia, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=47604, accessed on February 13, 2016.

Citation for the whole project: Jepson Flora Project (eds.) 2016. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on February 13, 2016.


Utricularia intermedia
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© 2004 Barry Rice
Utricularia intermedia
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© 2011 Barry Rice
Utricularia intermedia
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© 2006 Barry Rice
Utricularia intermedia
click for enlargement
© 2009 Barry Rice
Utricularia intermedia
click for enlargement
© 2004 Barry Rice
Utricularia intermedia
click for enlargement
© 2009 Barry Rice

More photos of Utricularia intermedia in CalPhotos


Geographic subdivisions for Utricularia intermedia:
CaR, SNH, MP;
Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.