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Ratibida columnifera

Higher Taxonomy
Family: Asteraceae (Compositae)View DescriptionDichotomous Key

Habit: Annual to tree. Leaf: basal and/or cauline, alternate, opposite, rarely whorled, simple to 2+ × compound. Inflorescence: 1° inflorescence a head, resembling a flower, of several types (see below), 1--many in generally +- cyme-like cluster; each head generally with +- calyx-like involucre of 1--many series of phyllaries (involucre bracts); receptacle of head flat to conic or columnar, paleate (bearing paleae = receptacle bracts) or epaleate; flowers 1--many per head. Flower: bisexual, unisexual, or sterile, +- small, of several types (see below); calyx 0 or modified into +- persistent pappus of bristles, scales, and/or awns; corolla radial or bilateral (0), lobes generally (0)3--5; stamens 4--5, filaments generally free, generally fused to corolla at tube/throat junction, anthers generally fused into cylinder around style, anther base generally rounded or cordate (deeply sagittate or with tail-like appendages), tip (= flattened appendage) generally projecting beyond pollen sac; pistil 1, 2-carpeled, ovary inferior, 1-chambered, 1-seeded, placenta basal, style 1, tip generally +- 2-branched (except in some staminate disk flowers), branch tips truncate or generally bearing +- brush-like appendages; stigmas 2, generally on adaxial faces of style branches. Fruit: achene (also called a cypsela) (drupe in Chrysanthemoides), cylindric to ovoid, sometimes compressed, generally deciduous with pappus attached.
Genera In Family: +- 1500 genera, 23000 species: worldwide, many habitats. Note: Flower and head types differ in form and sexual condition. A disk flower has a generally radial corolla, with a cylindric tube, expanded throat, and generally 5 lobes. Disk flowers are generally bisexual and fertile but occasionally staminate with reduced ovaries. Discoid heads comprise only disk flowers. A radiant head is a variant of a discoid head, with peripheral disk flower corollas expanded, often bilateral. A ray flower corolla is bilateral, generally with a slender tube and flattened petal-like ray (single lip composed of generally 3 lobes). Ray flowers are generally pistillate or sterile (occasionally lacking styles). Radiate heads have peripheral ray flowers and central disk flowers. Disciform heads superficially resemble discoid heads, with pistillate or sterile flowers that lack rays, together with or separate from disk flowers. A ligulate flower is bisexual, with a bilateral, generally ephemeral corolla and 5-lobed ligule. Liguliflorous heads comprise only ligulate flowers. See glossary p. 31 for illustrations of family characteristics. Echinops sphaerocephalus L., Gaillardia aristata Pursh, Gaillardia pulchella Foug., Hymenothrix loomisii S.F. Blake, Tagetes erecta L., Thelesperma megapotamicum (Spreng.) Kuntze are waifs. Melampodium perfoliatum Kunth, historic urban waif. Ageratum conyzoides L., Guizotia abyssinica (L. f.) Cass., Santolina chamaecyparisus L., orth. var. are rare or uncommon escapes from cultivation. Dyssodia papposa, Ismelia carinata (Schousb.) Sch. Bip. [Chrysanthemum carinatum Schousb.], Mantisalca salmantica (L.) Briq. & Cavill. are historical or extirpated waifs in California. Inula helenium L. not documented in California. Taxa of Aster in TJM (1993) treated here in Almutaster, Eucephalus, Eurybia, Ionactis, Oreostemma, Sericocarpus, Symphyotrichum; Chamomilla in Matricaria; Cnicus in Centaurea; Conyza in Erigeron and Laennecia; Dugaldia in Hymenoxys; Erechtites in Senecio; Hymenoclea in Ambrosia; Lembertia in Monolopia; Osteospermum ecklonis in Dimorphotheca; Picris echioides in Helminthotheca; Prionopsis in Grindelia; Raillardiopsis in Anisocarpus and Carlquistia; Schkuhria multiflora in Bahia; Trimorpha in Erigeron; Venidium in Arctotis; Whitneya in Arnica. Taxa of Arida in TJM2 (2012) treated here in Leucosyris.
Unabridged Note: Largest family of vascular plants in California and of eudicots globally.
eFlora Treatment Author: David J. Keil, except as noted
Scientific Editor: David J. Keil, Bruce G. Baldwin.
Genus: RatibidaView Description 

Habit: Perennial herb from taproot, rhizome, or caudex. Stem: 1--many from base, variously branched, generally erect, ribbed, strigose or rough-hairy. Leaf: basal and cauline, alternate, petioled, blade linear to ovate in outline, generally pinnately 1--2 × divided, the basal and distal-most sometimes entire; faces strigose to rough-hairy, gland-dotted. Inflorescence: heads radiate, 1 or in open, few-headed clusters; peduncle slender, bracts 0--few; involucre rotate; phyllaries in 2 unequal series, linear or narrowly triangular, spreading in flower, reflexed in fruit, outer generally longer; receptacle +- spheric to columnar, paleate; palea ovate, folded around fruit, tip hair-tufted, margins each with oblong resin gland. Ray Flower: 3--15+, sterile; corolla yellow, maroon, or 2-colored yellow and maroon; ray oblong to widely obovate. Disk Flower: 50--400+; corolla yellow-green or distally +- purple; anthers generally brown-purple; style tips triangular, long papillate. Fruit: oblanceolate, +- 4-angled, compressed side-to-side, winged, ciliate on abaxial margin; pappus 0 or 1--2 tooth-like projections or a low crown.
Species In Genus: 7 species: North America. Etymology: (Dacian name for a sp. of Aster)

Ratibida columnifera (Nutt.) Wooton & Standl.
Stem: 3--10 dm, finely strigose. Leaf: cauline divided into linear to oblong or narrowly elliptic, lobes; faces strigose. Inflorescence: heads 1 at stem tips and from distal leaf axils; peduncle 10--40 cm; receptacle cylindric. Ray Flower: 4--12; ray 7--30 mm, ovate to obovate, 2--3-lobed. Disk Flower: corolla 1--2.5 mm. Fruit: 2--3 mm; pappus of 0--2 minute teeth. Chromosomes: 2n=28.
Ecology: Dry, rocky slopes, upland valleys, open pine forest; Elevation: < 1600 m. Bioregional Distribution: PR, MP; Distribution Outside California: native North America prairies, British Columbia to Ontario, southern to Arizona, Louisiana, northern Mexico. Flowering Time: May--Aug Note: Incl in seed mixes; occasionally escapes or persists from cultivation.
Synonyms: Ratibida columnaris (Pursh) D. Don var. pulcherrima (DC.) D.J.N. Hind
eFlora Treatment Author: David J. Keil
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Citation for this treatment: David J. Keil 2016. Ratibida columnifera, in Jepson Flora Project (eds.) Jepson eFlora,, accessed on May 31, 2016.

Citation for the whole project: Jepson Flora Project (eds.) 2016. Jepson eFlora,, accessed on May 31, 2016.

Geographic subdivisions for Ratibida columnifera:
Markers link to CCH specimen records. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues. Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.