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Sherardia arvensis
FIELD MADDER

Higher Taxonomy
Family: RubiaceaeView DescriptionDichotomous Key

Common Name: MADDER FAMILY
Habit: Annual to tree, vine. Leaf: generally opposite (whorled), entire; stipules generally fused to stem, adjacent pairs occasionally fused, or occasionally leaf-like and appearing like whorled leaves. Inflorescence: cyme, panicle, spike, cluster, or flower 1, generally terminal and +- axillary. Flower: generally bisexual; calyx +- 4(5)-lobed, occasionally 0 (Galium, Crucianella) or 6 (Sherardia); corolla generally radial, 4(5)-lobed; stamens epipetalous, alternate corolla lobes, generally included; ovary generally inferior, chambers generally 2 or 4, style 1(2). Fruit: drupe, berry, or 2 or 4 nutlets [capsule].
Genera In Family: +- 500 genera, 6000 species: worldwide, especially tropics; many cultivated, including Coffea, coffee; Cinchona, quinine; many ornamental. Note: Diodia teres Walter doubtfully in California.
eFlora Treatment Author: Robert E. Preston & Lauramay T. Dempster, except as noted
Scientific Editor: Douglas H. Goldman, Bruce G. Baldwin.
Genus: SherardiaView Description 


Species In Genus: 1 sp. Etymology: (Wm. Sherard, Dillenius' patron, John Ray's friend, 1659--1728)

Sherardia arvensis L.
NATURALIZED
Habit: Annual, matted. Stem: decumbent, generally many-branched at base, 7--16 cm, 4-angled. Leaf: whorls of 5--6; 4--13 mm, lanceolate or oblanceolate, margin thick, tip acute or weak-spined, stipules leaf-like. Inflorescence: head, axillary, involucred; flowers sessile, generally 2--3, generally included in involucre; involucre bracts +- free, +- = leaves, generally < peduncle. Flower: calyx lobes 6, persistent, tips +- dissected; corolla salverform, generally 4-lobed, pink or lavender; styles 2, thread-like, fused proximally. Fruit: 2 nutlets; hairs soft. Chromosomes: 2n=22.
Ecology: Pastures, disturbed areas, grassland, dry meadows, oak woodland; Elevation: 10--1160 m. Bioregional Distribution: NCo, NCoRO, NCoRI, CaRF, n SN, ScV, SnFrB, SCoRO, SCo; Distribution Outside California: to Idaho, Arizona, eastern United States; native to Mediterranean. Flowering Time: Mar--Jul
eFlora Treatment Author: Robert E. Preston & Lauramay T. Dempster
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Citation for this treatment: Robert E. Preston & Lauramay T. Dempster 2016. Sherardia arvensis, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=44299, accessed on February 07, 2016.

Citation for the whole project: Jepson Flora Project (eds.) 2016. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on February 07, 2016.


Sherardia arvensis
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© 2008 Steve Matson
Sherardia arvensis
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© 2008 Steve Matson
Sherardia arvensis
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© 2004 Carol W. Witham
Sherardia arvensis
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© 2009 Julie Kierstead Nelson
Sherardia arvensis
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© 2005 Steve Matson
Sherardia arvensis
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© 2008 Keir Morse

More photos of Sherardia arvensis in CalPhotos


Geographic subdivisions for Sherardia arvensis:
NCo, NCoRO, NCoRI, CaRF, n SN, ScV, SnFrB, SCoRO, SCo;
Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.