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Pyrola aphylla
LEAFLESS WINTERGREEN


Higher Taxonomy
Family: EricaceaeView DescriptionDichotomous Key
Common Name: HEATH FAMILY
Habit: Perennial herb, shrub, tree. Stem: bark often peeling distinctively. Leaf: simple or 0, generally cauline, alternate, opposite (whorled), evergreen or deciduous, often leathery, petioled or not; stipules 0. Inflorescence: raceme, panicle, cyme, or flowers 1, terminal or axillary, generally bracted; pedicel often with 2 bractlets. Flower: generally bisexual, generally radial, bell-shaped, cylindric, or urn-shaped; sepals generally (0)4--5, generally free; petals generally (0)4--5, free or fused; stamens (2--5)8--10, free, filaments rarely appendaged, anthers dehiscing by pores or slits, awns 0 or 2(4), seemingly abaxial, reduced or elongate, generally curved; nectary generally present at ovary base, generally disk-like; ovary superior or inferior, chambers generally 1--5, placentas axile or parietal, ovules 1--many per chamber, style 1, stigma head- to funnel-like or lobed. Fruit: capsule, drupe, berry. Seed: generally many, winged or not.
Genera In Family: +- 100 genera, 3000 species: generally worldwide except deserts; some cultivated, especially Arbutus, Arctostaphylos, Rhododendron, Vaccinium. Note: Monophyletic only if Empetraceae included, as treated here. Ledum included in Rhododendron. Non-green plants obtain nutrition from green plants through fungal intermediates.
eFlora Treatment Author: Gary D. Wallace, except as noted
Scientific Editor: Gary D. Wallace, Thomas J. Rosatti, Bruce G. Baldwin.
Genus: PyrolaView DescriptionDichotomous Key


Common Name: WINTERGREEN
Habit: Perennial subshrub, evergreen, rhizomed. Leaf: +- basal, reniform, ovate, +- round, elliptic, or obovate, +- entire to crenate or dentate, generally petioled (if vestigial, lanceolate to deltate, +- not green, sessile). Inflorescence: raceme, +- erect, not 1-sided, elongate; scape smooth, glabrous, bracts generally 1--several, deltate or lanceolate; flower bract 1 per flower, < to >> pedicel. Flower: radial, +- closed or subradial or bilateral, +- open, parts in 5s, free; petals without tubercles, upper 2 generally forming hood over upturned stamens; stamens 10, filaments generally widened at base, smooth, glabrous, anther generally with tube-like constrictions subtending pores; nectary 0; ovary superior, style +- included and straight or generally exserted and downcurved, stigma peltate, with 5 spreading lobes above a prominent, reflexed collar or generally not peltate, generally with 5 +- erect lobes projecting beyond a delicate, reflexed collar. Fruit: capsule, pendent; valves opening base to tip, margins fibrous. Seed: +- 1000.
Species In Genus: +- 30 species: generally circumboreal, high mountains of Central America, Sumatra. Etymology: (Latin: little pear, +- from leaf shape) Note: Pyrola chlorantha not in CA.
Unabridged Note: In Jepson eFlora Revision 2, Pyrola aphylla, Pyrola crypta, Pyrola dentata added, as native.
eFlora Treatment Author: Diana D. Jolles
Reference: Jolles & Wolfe 2012 Syst Bot 37:468--477; Jolles & Wilson 2014 Taxon 63(4):789--800
Pyrola aphylla Sm.
NATIVE
Leaf: blade generally vestigial or 0, largest +-1 cm, lanceolate to deltate, +- not green, adaxially dull, occasionally with +- light-color along adaxial veins, often hidden by leaf litter or loose substrate; petiole 0. Inflorescence: < 6 dm including scape, often several together; flowers 7--25; flower bract 0.5--1.5 × pedicel. Flower: bilateral, +- open; sepals generally 2--3 mm; petals 5--7.5 mm, cream-white or pink; anthers 2.8--4.5 mm. Chromosomes: 2n=46.
Ecology: Mixed conifer forest to Quercus, Arbutus, or Pinus woodland, forested serpentine areas, in deep litter and duff on hillsides of decomposed granite or loose, coarse sand; Elevation: 500--2500 m. Bioregional Distribution: NW, CaR, SNF, SNH, CW, SnBr, PR, MP; Distribution Outside California: to southwest British Columbia, Baja California. Flowering Time: Jun--Aug Note: Occasionally hybridizes with other species and produces clones with blades expanded to +- 1 cm wide.
Unabridged Synonyms: Pyrola picta f. aphylla (Sm.) Camp, Thelaia aphylla (Sm.) Alef.
Jepson eFlora Author: Diana D. Jolles
Reference: Jolles & Wolfe 2012 Syst Bot 37:468--477; Jolles & Wilson 2014 Taxon 63(4):789--800
Unabridged Reference: Camp 1940 Bull Torrey Bot Club 67:453--465; Haber 1987 Syst Bot 12:324--335
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Citation for this treatment: Diana D. Jolles 2017, Pyrola aphylla, in Jepson Flora Project (eds.) Jepson eFlora, Revision 5, http://ucjeps.berkeley.edu/eflora/eflora_display.php?tid=40453, accessed on October 15, 2019.

Citation for the whole project: Jepson Flora Project (eds.) 2019, Jepson eFlora, http://ucjeps.berkeley.edu/eflora/, accessed on October 15, 2019.

Pyrola aphylla
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© 2009 Thomas Stoughton
Pyrola aphylla
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© 2012 Barry Rice
Pyrola aphylla
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© 2009 Thomas Stoughton
Pyrola aphylla
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© 2009 Thomas Stoughton
Pyrola aphylla
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© 2010 Keir Morse
Pyrola aphylla
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© 2010 Keir Morse

More photos of Pyrola aphylla in CalPhotos



Geographic subdivisions for Pyrola aphylla:
NW, CaR, SNF, SNH, CW, SnBr, PR, MP
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).





 

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Data provided by the participants of the  Consortium of California Herbaria.
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All markers link to CCH specimen records. The original determination is shown in the popup window.
Blue markers indicate specimens that map to one of the expected Jepson geographic subdivisions (see left map). Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa, if there are more than 1 infraspecific taxon in CA.
Blue line denotes eFlora flowering time.