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Lagophylla ramosissima

Higher Taxonomy
Family: Asteraceae (Compositae)View DescriptionDichotomous Key

Habit: Annual to tree. Leaf: basal and/or cauline, alternate, opposite, rarely whorled, simple to 2+ × compound. Inflorescence: 1° inflorescence a head, resembling a flower, of several types (see below), 1--many in generally +- cyme-like cluster; each head generally with +- calyx-like involucre of 1--many series of phyllaries (involucre bracts); receptacle of head flat to conic or columnar, paleate (bearing paleae = receptacle bracts) or epaleate; flowers 1--many per head. Flower: bisexual, unisexual, or sterile, +- small, of several types (see below); calyx 0 or modified into +- persistent pappus of bristles, scales, and/or awns; corolla radial or bilateral (0), lobes generally (0)3--5; stamens 4--5, filaments generally free, generally fused to corolla at tube/throat junction, anthers generally fused into cylinder around style, anther base generally rounded or cordate (deeply sagittate or with tail-like appendages), tip (= flattened appendage) generally projecting beyond pollen sac; pistil 1, 2-carpeled, ovary inferior, 1-chambered, 1-seeded, placenta basal, style 1, tip generally +- 2-branched (except in some staminate disk flowers), branch tips truncate or generally bearing +- brush-like appendages; stigmas 2, generally on adaxial faces of style branches. Fruit: achene (also called a cypsela) (drupe in Chrysanthemoides), cylindric to ovoid, sometimes compressed, generally deciduous with pappus attached.
Genera In Family: +- 1500 genera, 23000 species: worldwide, many habitats. Note: Flower and head types differ in form and sexual condition. A disk flower has a generally radial corolla, with a cylindric tube, expanded throat, and generally 5 lobes. Disk flowers are generally bisexual and fertile but occasionally staminate with reduced ovaries. Discoid heads comprise only disk flowers. A radiant head is a variant of a discoid head, with peripheral disk flower corollas expanded, often bilateral. A ray flower corolla is bilateral, generally with a slender tube and flattened petal-like ray (single lip composed of generally 3 lobes). Ray flowers are generally pistillate or sterile (occasionally lacking styles). Radiate heads have peripheral ray flowers and central disk flowers. Disciform heads superficially resemble discoid heads, with pistillate or sterile flowers that lack rays, together with or separate from disk flowers. A ligulate flower is bisexual, with a bilateral, generally ephemeral corolla and 5-lobed ligule. Liguliflorous heads comprise only ligulate flowers. See glossary p. 31 for illustrations of family characteristics. Echinops sphaerocephalus L., Gaillardia aristata Pursh, Gaillardia pulchella Foug., Hymenothrix loomisii S.F. Blake, Tagetes erecta L., Thelesperma megapotamicum (Spreng.) Kuntze are waifs. Melampodium perfoliatum Kunth, historic urban waif. Ageratum conyzoides L., Guizotia abyssinica (L. f.) Cass., Santolina chamaecyparisus L., orth. var. are rare or uncommon escapes from cultivation. Dyssodia papposa, Ismelia carinata (Schousb.) Sch. Bip. [Chrysanthemum carinatum Schousb.], Mantisalca salmantica (L.) Briq. & Cavill. are historical or extirpated waifs in California. Inula helenium L. not documented in California. Taxa of Aster in TJM (1993) treated here in Almutaster, Eucephalus, Eurybia, Ionactis, Oreostemma, Sericocarpus, Symphyotrichum; Chamomilla in Matricaria; Bahia in Hymenothrix; Cnicus in Centaurea; Conyza in Erigeron and Laennecia; Dugaldia in Hymenoxys; Erechtites in Senecio; Hymenoclea in Ambrosia; Lembertia in Monolopia; Osteospermum ecklonis in Dimorphotheca; Picris echioides in Helminthotheca; Prionopsis in Grindelia; Raillardiopsis in Anisocarpus and Carlquistia; Schkuhria multiflora in Picradeniopsis; Trimorpha in Erigeron; Venidium in Arctotis; Whitneya in Arnica. Amauriopsis in TJM2 (2012) treated here in Hymenothrix; Arida in Leucosyris; Bahia in Picradeniopsis.
Unabridged Note: Largest family of vascular plants in California and of eudicots globally.
eFlora Treatment Author: David J. Keil, except as noted
Scientific Editor: David J. Keil, Bruce G. Baldwin.
Genus: LagophyllaView DescriptionDichotomous Key

Common Name: HARE-LEAF
Habit: Annual 1--10(15) dm. Stem: +- erect. Leaf: mostly cauline, proximal opposite, most alternate, +- sessile; blade narrowly elliptic to linear or proximal oblanceolate to spoon-shaped, entire or proximal sometimes toothed, coarse-, soft-, or silky-hairy or strigose, all or distal sometimes also stalked-glandular. Inflorescence: heads radiate, in tight groups or +- panicle-like clusters; involucre 3--6+ mm diam, +- hemispheric or obovoid to obconic, sometimes subtended by calyx-like set of bracts; phyllaries 5 in 1 series, linear to oblanceolate, each wholly enveloping a subtended ray ovary, spreading and falling with fruit, on angles coarsely long-straight-hairy to minutely coarse-hairy or scabrous; receptacle flat to convex, densely bristly; paleae in ring between ray and disk flowers, fused or free, scarious. Ray Flower: 5; corolla yellow, ray fan-shaped, deeply lobed, abaxially often red- to purple-veined. Disk Flower: 6, staminate; corolla yellow, tube < throat, lobes deltate; anthers +- dark purple, tips triangular-ovate to +- rounded; style glabrous proximal to undivided, awl-shaped, short-hairy tip. Fruit: ray fruit +- compressed front-to-back, glabrous, black, tip beakless, pappus 0; disk fruit 0, pappus 0.
Species In Genus: 5 species: California, to Washington, Montana, Nevada. Etymology: (Greek: hare leaf, for soft-hairy leaves) Note: Easily overlooked; leaves wither early, heads close at mid-day. Lagophylla diabolensis newly described, added as native.
eFlora Treatment Author: Bruce G. Baldwin
Reference: Baldwin 2013 Madroño 60:249--254
Unabridged Reference: Thompson 1983 Ph.D. Dissertation, Univ of California, Davis; Baldwin & Strother 2006 FNANM 21:260--261

Lagophylla ramosissima Nutt.
Habit: Plant 1--10(15) dm; self-compatible. Stem: glandless; main axis obvious or not (+- zigzag). Leaf: +- gray, most glandless, distal stalked-glandular abaxially, glands generally +- white or yellow, sometimes golden-brown. Inflorescence: heads tightly grouped or in open, panicle-like clusters; calyx-like bracts subtending involucre 2--5; involucre obconic to obovoid; phyllaries 4--7 mm, on angles coarsely long-spreading-hairy, hairs +- widely spreading to curved toward phyllary tip, 0.5--1+ mm. Ray Flower: ray 3--6 mm. Fruit: dull to +- shiny, weakly striate. Chromosomes: 2n=14.
Ecology: Grassland, openings in scrub, woodland, forest; Elevation: < 1800 m. Bioregional Distribution: CA-FP (exc NCo, ChI, SnJt), MP; Distribution Outside California: to Washington, Montana, Nevada. Flowering Time: Apr--Oct
Synonyms: Lagophylla ramosissima subsp. congesta (Greene) D.D. Keck; Lagophylla ramosissima Nutt. subsp. ramosissima;
Jepson eFlora Author: Bruce G. Baldwin
Reference: Baldwin 2013 Madroño 60:249--254
Unabridged Reference: Thompson 1983 Ph.D. Dissertation, Univ of California, Davis; Baldwin & Strother 2006 FNANM 21:260--261
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botanical illustration including Lagophylla ramosissima


Citation for this treatment: Bruce G. Baldwin 2014, Lagophylla ramosissima, in Jepson Flora Project (eds.) Jepson eFlora, Revision 2,, accessed on November 17, 2018.

Citation for the whole project: Jepson Flora Project (eds.) 2018, Jepson eFlora,, accessed on November 17, 2018.

Lagophylla ramosissima
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© 2005 Michael Charters
Lagophylla ramosissima
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© 2010 Neal Kramer
Lagophylla ramosissima subsp. ramosissima
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© 2008 Steve Matson
Lagophylla ramosissima
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© 2016 Neal Kramer
Lagophylla ramosissima
click for enlargement
© 2010 Neal Kramer
Lagophylla ramosissima
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© 2016 Neal Kramer

More photos of Lagophylla ramosissima in CalPhotos

Geographic subdivisions for Lagophylla ramosissima:
CA-FP (exc NCo, ChI, SnJt), MP;
Markers link to CCH specimen records. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues. Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa, if there are more than 1 infraspecific taxon in CA.
Blue line denotes eFlora flowering time.