Jepson eFlora: Taxon page
Vascular Plants of California
Key to families | Table of families and genera
Previous taxon Index to accepted names and synonyms:
| A | B | C | D | E | F | G | H | I | J | K | L | M | N | O | P | Q | R | S | T | U | V | W | X | Y | Z |
Next taxon


Pellaea breweri
BREWER'S CLIFF-BRAKE


Higher Taxonomy
Family: PteridaceaeView DescriptionDichotomous Key
Common Name: BRAKE FAMILY
Habit: Perennial herb, in soil or on or among rocks; rhizome creeping to erect, scaly. Leaf: generally all +- alike (or of 2 kinds, fertile, sterile), generally < 50 cm, often < 25 cm; stipe generally thin, wiry, often dark, ×-section with vascular strands generally 1--3, less often many in circle; blade generally pinnate or +- palmate-pinnate (see Adiantum), often >= 2-compound, abaxially often with glands, +- powdery exudate, hairs, or scales; segments round, oblong, fan-shaped, or other, veins generally free. Sporangia: in sori or not, marginal, submarginal, or along veins, covered by recurved, often modified segment margins (false indusia) or not; true indusia 0; spores spheric, sides flat or not, scar with 3 radiating branches.
Genera In Family: +- 40 genera, 500 species: worldwide, especially dry areas. Note: CA members of Cheilanthes moved to the distantly related Myriopteris; Pellaea breweri to be moved as well, from a to-be-redefined Pellaea; traditional, often untenable limits of genera outside CA also being clarified using molecular phylogenetics.
eFlora Treatment Author: Ruth E.B. Kirkpatrick, Alan R. Smith & Thomas Lemieux, except as noted
Scientific Editor: Alan R. Smith, Thomas J. Rosatti.
Genus: PellaeaView DescriptionDichotomous Key


Common Name: CLIFF-BRAKE
Habit: Plant in soil or rock crevices; rhizome short- to long-creeping, scales overlapped, narrowly linear, light- to red- or medium-brown, often with dark mid-stripe. Leaf: erect, persistent, < 1 m; stipes +- cylindric, generally dark or red-brown to +- black, +- shiny, glabrous; blade 1--4-pinnate; segments generally stalked, generally free, linear to rounded, lobed or not, often folded lengthwise when dried; veins generally free. Sporangia: in +- continuous, submarginal bands, among a +- white to +- yellow exudate or not; segment margin generally recurved, generally modified; spores tan to light yellow.
Species In Genus: +- 35 species: tropics, temperate, few in Europe, 0 in Asia. Etymology: (Greek: dusky, from blue-gray leaves) Note: Occasionally cultivated; as defined by Tryon (1957), polyphyletic (Kirkpatrick, 2007).
Unabridged Note: Occasionally cultivated. Molecular data suggest Pellaea in the sense of Tryon (1957) is polyphyletic with monophyletic Astrolepis and Pellaea sect. Platyloma, as well as elements of Paragymnopteris and Paraceterach nested within (Kirkpatrick, 2007).
eFlora Treatment Author: Ruth E.B. Kirkpatrick, Alan R. Smith, Thomas Lemieux & Edward Alverson
Reference: Kirkpatrick 2007 Syst Bot 32:504--518
Unabridged Reference: Kirkpatrick, R.E.B. 2007. Investigating the monophyly of Pellaea (Pteridaceae) in the context of a phylogenetic analysis of cheilanthoid ferns. Syst Bot 32:504--518; Tryon 1957 Ann Missouri Bot Gard 44(2):125--193
Pellaea breweri D.C. Eaton
NATIVE
Habit: Rhizome short-creeping, branched, > 10 cm, 5(7) mm wide; scales +- thread-like, red-brown, dark mid-stripe 0. Leaf: clustered, 8--20(25) cm, 2--3(4) cm wide, pale greenish; stipe < 2 mm wide, fracture lines at base many; blade 1-pinnate, oblong, rachis green at tip; pinnae < 2 cm, < 1.5 cm wide, lance-ovate, deeply 2(3)-lobed. Sporangia: 64-spored; spores dark to light brown. Chromosomes: 2n=58.
Ecology: Generally n-facing granite rock crevices, slopes; Elevation: 1500--3700 m. Bioregional Distribution: KR, CaRH, SNH, SnBr, GB, DMtns; Distribution Outside California: to Washington, Idaho, Colorado.
Unabridged Note: Molecular evidence suggests that Paragymnopteris marantae (L.) K.H. Shing of Eurasia and Paragymnopteris sargentii (H. Christ) K.H. Shing of China are sister to each other, that both are sister to Pellaea breweri, and that these 3 form the earliest diverging pellaeoid clade (Kirkpatrick 2007).
Jepson eFlora Author: Ruth E.B. Kirkpatrick, Alan R. Smith, Thomas Lemieux & Edward Alverson
Reference: Kirkpatrick 2007 Syst Bot 32:504--518
Index of California Plant Names (ICPN; linked via the Jepson Online Interchange)

Previous taxon: Pellaea brachyptera
Next taxon: Pellaea bridgesii

Name Search

Botanical illustration including Pellaea breweri

botanical illustration including Pellaea breweri

Contact/Feedback

Citation for this treatment: Ruth E.B. Kirkpatrick, Alan R. Smith, Thomas Lemieux & Edward Alverson 2012, Pellaea breweri, in Jepson Flora Project (eds.) Jepson eFlora, eflora_display.php?tid=36778, accessed on January 21, 2020.

Citation for the whole project: Jepson Flora Project (eds.) 2020, Jepson eFlora, http://ucjeps.berkeley.edu/eflora/, accessed on January 21, 2020.

Pellaea breweri
click for enlargement
© 2004 Larry Blakely
Pellaea breweri
click for enlargement
© 2004 Larry Blakely
Pellaea breweri
click for enlargement
© 2011 Barry Breckling
Pellaea breweri
click for enlargement
© 1998 Larry Blakely
Pellaea breweri
click for enlargement
© 2005 Dieter Wilken
Pellaea breweri
click for enlargement
© 2004 Larry Blakely

More photos of Pellaea breweri in CalPhotos



Geographic subdivisions for Pellaea breweri:
KR, CaRH, SNH, SnBr, GB, DMtns
MAP CONTROLS
1. You can change the display of the base map and layers by clicking on the layer control box in the upper right-hand corner.
2. California county polygons can be turned off and on in the layer control box.
3. Filling of Jepson subdivision polygons can be turned off and on in the layer control box.
4. Moving the cursor over any numbered cluster will show the range boundary of the included specimens (with a blue polygon).
5. Marker clustering can be turned off by clicking this link:      Marker Clustering OFF
WARNING: Turning this off might cause maps with large numbers of specimens to load slowly.
map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).





 

View elevation by latitude chart


Data provided by the participants of the  Consortium of California Herbaria.
MAP LEGEND
View all CCH records
All markers link to CCH specimen records. The original determination is shown in the popup window.
Blue markers indicate specimens that map to one of the expected Jepson geographic subdivisions (see left map). Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
READ ABOUT YELLOW FLAGS


CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa, if there are more than 1 infraspecific taxon in CA.
Blue line denotes eFlora flowering time (fruiting time in some monocot genera).