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Pectocarya heterocarpa
MIXED-NUT PECTOCARYA


Higher Taxonomy
Family: BoraginaceaeView DescriptionDichotomous Key
Common Name: BORAGE or WATERLEAF FAMILY
Habit: Annual to shrub or small tree, or non-green root parasite, often bristly or sharp-hairy. Stem: prostrate to erect. Leaf: basal and/or cauline, generally simple, generally alternate. Inflorescence: generally cymes, or panicle-, raceme-, head-, or spike-like, generally coiled in flower (often described as scorpioid), generally elongating in fruit, or flowers 1--2 per axil. Flower: bisexual, generally radial; sepals (4)5(10), fused at least at base, or free; corolla (4)5(10)-lobed, salverform, funnel-shaped, rotate, or bell-shaped, generally without scales at tube base, with 0 or 5 appendages at tube top, alternate stamens; stamens epipetalous; ovary generally superior, entire to 4-lobed, style 1(2), entire or 2-lobed or -branched. Fruit: valvate or circumscissile capsule or nutlets 1--4, free (fused), smooth to roughened, prickly or bristly or not.
Genera In Family: +- 120 genera, +- 2300 species: tropics, temperate, especially western North America, Mediterranean; some cultivated (Borago, Heliotropium, Echium, Myosotis, Nemophila, Phacelia, Symphytum, Wigandia). Toxicity: Many genera may be TOXIC from pyrrolizidine alkaloids or accumulated nitrates. Note: Recently treated to include Hydrophyllaceae, Lennoaceae. Wigandia urens added, as naturalized.
eFlora Treatment Author: Ronald B. Kelley, Robert Patterson, Richard R. Halse & Timothy C. Messick, family description, key to genera, treatment of genera by Ronald B. Kelley, except as noted
Scientific Editor: Ronald B. Kelley, Robert Patterson, Thomas J. Rosatti, Bruce G. Baldwin, David J. Keil.
Genus: PectocaryaView DescriptionDichotomous Key


Common Name: PECTOCARYA, COMBSEED
Habit: Annual. Stem: 2--40 cm, strigose, breaking at nodes or not. Leaf: generally alternate, generally 0.5--4 cm, +- linear, strigose to sharp-bristled. Inflorescence: bracted or not; pedicel in fruit generally free from nutlets, generally recurved. Flower: basal cleistogamous flowers generally 0; calyx generally < fruit, lobes free, not arched over 1 nutlet, with hooked or straight prickles, in fruit +- equal or, if unequal, upper 2 > others; corolla funnel-shaped, white, limb 0.5--2.5 mm diam, appendages white or yellow; style attached to receptacle, unbranched, generally persistent, stigma 1, head-like. Fruit: nutlets generally 4, generally paired, often dissimilar in shape, ornamentation, margin width, spreading, 1--4.5 mm, generally compressed, marginal prickles straight or hooked at tip.
Species In Genus: 15 species: to British Columbia, Wyoming, Texas, northwestern Mexico; South America. Etymology: (Greek: comb nut, from bristly to dentate nutlet margins of some species) Note: Nutlets of basal, cleistogamous flowers (present in Pectocarya heterocarpa, Pectocarya peninsularis) unusual, not to be used in key.
Unabridged Note: Flowers generally of 1 kind with observable white corolla occurring on the cauline inflorescence sections, except for Pectocarya heterocarpa, Pectocarya peninsularis, which also produce cleistogamous flowers, without an observable corolla, that occur at stem bases and produce nutlet shapes not represented in the diagnostic key.
eFlora Treatment Author: Ronald B. Kelley
Unabridged Reference: Veno 1979 Ph.D. Dissertation Univ of California, Los Angeles
Pectocarya heterocarpa (I.M. Johnst.) I.M. Johnst.
NATIVE
Stem: prostrate to ascending, 2--25 cm. Inflorescence: pedicel in fruit 1.3--2.8 mm, partly fused to 1 nutlet. Flower: basal cleistogamous flowers present; lower 3 calyx lobes in fruit unequal, < others. Fruit: nutlets 1.2--3 mm, oblanceolate to oblong; cauline paired, curved in 2 planes, 1 pair incurved, the other pair recurved, margins +- entire, narrow- to wide-membranous, to dentate. Chromosomes: 2n=24.
Ecology: Washes, roadsides, openings in creosote-bush scrub, Joshua-tree woodland; Elevation: < 1600 m. Bioregional Distribution: s SNF, Teh, SW, W&I, D; Distribution Outside California: to southwestern Utah, western Texas, northern Mexico. Flowering Time: Feb--May Note: Basal nutlets 2--4 per flower, not paired, reflexed, 1 unmargined, others narrow-entire- to +- -dentate-margined.
Jepson eFlora Author: Ronald B. Kelley
Unabridged Reference: Veno 1979 Ph.D. Dissertation Univ of California, Los Angeles
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botanical illustration including Pectocarya heterocarpa

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Citation for this treatment: Ronald B. Kelley 2012, Pectocarya heterocarpa, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/eflora/eflora_display.php?tid=36528, accessed on June 19, 2019.

Citation for the whole project: Jepson Flora Project (eds.) 2019, Jepson eFlora, http://ucjeps.berkeley.edu/eflora/, accessed on June 19, 2019.

Pectocarya heterocarpa
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© 2010 Neal Kramer
Pectocarya heterocarpa
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© 2006 Steve Matson
Pectocarya heterocarpa
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© 2011 Steve Matson
Pectocarya heterocarpa
click for enlargement
© 2011 Steve Matson
Pectocarya heterocarpa
click for enlargement
© 2006 Steve Matson
Pectocarya heterocarpa
click for enlargement
© 2011 Steve Matson

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Geographic subdivisions for Pectocarya heterocarpa:
s SNF, Teh, SW, W&I, D
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).





 

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All markers link to CCH specimen records. The original determination is shown in the popup window.
Blue markers indicate specimens that map to one of the expected Jepson geographic subdivisions (see left map). Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa, if there are more than 1 infraspecific taxon in CA.
Blue line denotes eFlora flowering time.